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R1a has lower frequencies in south-eastern Europe than in north-eastern Europe but at the same time unusual STR diversity exist among Balkan R1a.
I2-dinaric on the other hand has higher frequencies in European south-east than elsewhere in Europe but much lower diversity.
Historically, no other part of Europe was invaded a higher number of times by steppe peoples than the Balkans. Chronologically, the first R1a invaders came with the westward expansion of the Yamna culture (from 4200 BCE), a succession of steppe migrations that lasted about 2000 years. Then came the Thracians (1500 BCE), followed by the Illyrians (around 1200 BCE), the Huns and the Alans (400 CE), the Avars, the Bulgars and the Serbs (all around 600 CE), and the Magyars (900 CE), among others. These peoples originated from different parts of the Eurasian steppes, anywhere between Eastern Europe and Central Asia, which is why such high STR diversity is found within Balkanic R1a nowadays. It is not yet possible to determine the ethnic origin for each variety of R1a, apart from the fact that about any R1a is associated with tribes from Eurasian steppe at one point in history.What is the explanation for this ? Does it mean R1a was brought to the region by various groups from ancient Greeks, Thraco-Illyrians, Turkic Bulgars, Avars, and finally Slavs ?Localization of the ancestral haplogroup R1a1 is still controversial. Presumably, it originated in Eurasia. Today, there are two centers of its highest concentration - in North India and the Ukraine, as well as in the south of Russia. On the one hand, the high frequency of occurrence can be observed in any region with a large population, on the other hand its highest diversity is observed in the Balkans.
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