New paper! DNA of various Bronze Age Eurasian populations!

[Longbowman]

http://www.nature.com/nature/journal/v522/n7555/full/nature14507.html

[Graham]

The folk at Eurogenes seemed to be exited. Havent had a look, should be interesting cheers for posting it here.

[Dr. Robotnik the Subbotnik]

looks like you have to pay money to read more

[Jackson]

Looks to be really interesting, the files have been early released so expect a lot of activity over the next week i guess. :D

[Longbowman]

E3b 30% in Armenia. Just E3b and R1b (only Middle Eastern population).

http://www.nature.com/nature/journal/v522/n7555/images/nature14507-sf6.jpg

Seems that J rampaged through the Middle East in the Iron Age.

[Hevo]

Genetic structure of ancient Europe and the Pontic-Caspian steppe.
http://i58.tinypic.com/330428l.jpg

Fst distances.

http://i59.tinypic.com/16jeqkj.jpg

[Gaston]

E3b 30% in Armenia. Just E3b and R1b (only Middle Eastern population).

http://www.nature.com/nature/journal/v522/n7555/images/nature14507-sf6.jpg

Seems that J rampaged through the Middle East in the Iron Age.

Yeah, that's what struck me first. So E1b1b1 and R1b (xM269) in the Bronze age Middle East (with E1b being older in the region than R1b), followed by dramatic increase of J (or influx?) in the Iron Age?


EDIT: R1b-M269 might very well be present in the Bronze Age in the Near East but not during the Neolithic (as a compagnon of E1b).

[Longbowman]

Yeah, that's what struck me first. So E1b1b1 and R1b (xM269) in the Bronze age Middle East (with E1b being older in the region than R1b), followed by dramatic increase of J (or influx?) in the Iron Age?

That's what it seems like. But the rest of the Fertile Crescent needs to stop fighting and let us harvest DNA samples before we know for sure.

[DarknessInside]

No free full text of this study?

[Graham]

..

[Black Wolf]

E3b 30% in Armenia. Just E3b and R1b (only Middle Eastern population).

http://www.nature.com/nature/journal/v522/n7555/images/nature14507-sf6.jpg

Seems that J rampaged through the Middle East in the Iron Age.

I find it even more interesting that J2 and J2a have shown up in Iron Age Russia and Iron Age Altai.

[safinator]

http://www.nature.com/nature/journal/v522/n7555/images/nature14507-f2.jpg


Bronze Age Montenegrin samples are close to Tuscans and who's the most Tuscan like population in the Balkans?

:laugh:

[curupira]

The Bronze Age of Eurasia (around 3000–1000 BC) was a period of major cultural changes. However, there is debate about whether these changes resulted from the circulation of ideas or from human migrations, potentially also facilitating the spread of languages and certain phenotypic traits. We investigated this by using new, improved methods to sequence low-coverage genomes from 101 ancient humans from across Eurasia. We show that the Bronze Age was a highly dynamic period involving large-scale population migrations and replacements, responsible for shaping major parts of present-day demographic structure in both Europe and Asia. Our findings are consistent with the hypothesized spread of Indo-European languages during the Early Bronze Age. We also demonstrate that light skin pigmentation in Europeans was already present at high frequency in the Bronze Age, but not lactose tolerance, indicating a more recent onset of positive selection on lactose tolerance than previously thought.
http://www.nature.com/nature/journal/v522/n7555/full/nature14507.html

http://abload.de/img/nature14507-sf61rwr50.jpg

[Longbowman]

The ancient Armenian E3b has been shown to be my clade (E-M123). Autochtonous.

[curupira]

Very interesting. Afanasievo is significantly related to Yamnaya. And Remedello, long thought of as IE derived, is quite Neolithic European (with 0 R1a and 0 R1b). I had opened a thread on it here (one could delete the thread I opened or merge it with this one):

http://www.theapricity.com/forum/showthread.php?173706-Bronze-Age-Eurasians-(Allentoft-et-al-2015)&p=3633164#post3633164

[curupira]

“Now we see the Yamnaya is not only spreading north into Europe; they’re also spreading east, crossing the Urals, getting all the way into central Asia, all the way into the Altai, between Mongolia, China, and Siberia,” says evolutionary biologist Eske Willerslev of the University of Copenhagen...

This eastern branch of the Yamnaya (or Afanasievo) persisted in central Asia and, perhaps, Mongolia and China until they themselves were replaced by fierce warriors in chariots called the Sintashta (also known as the Andronovo culture). These people from the Urals and Caucuses, who were genetically related to central Europeans, persisted in central Asia until 2000 years ago, which means that people in central Asia were actually more like Europeans than living Asians. It wasn’t until relatively recently—just 2000 years ago—that these “Caucasians” were replaced by immigrants from eastern Asia, such as the Karasuk, Mezhovskaya, and other Iron Age cultures that today make up the ancestry of people in central Asia...

“There is a real sense that after more than 2 centuries of linguistics trying to solve the Indo-European question, it's ancient DNA that is suddenly moving us fast toward a possible resolution,” says linguist Paul Heggarty of the Max Planck Institute for Evolutionary Anthropology in Leipzig, Germany. But it is not enough just to have data from northern Eurasia, where the Yamnaya’s movements may reflect only one part of the spread of Indo-European languages. Heggarty adds: “We need key data from the majority of the Indo-European-speaking world in the Mediterranean and south of the Black Sea-Caspian-Himalayas.”
http://news.sciencemag.org/archaeology/2015/06/nomadic-herders-left-strong-genetic-mark-europeans-and-asians

[Hevo]

a Bell Beaker man from Osterhofen-Altenmarkt Germany turned out to be R1b U152.



Here is a list of ancient Arm0 Y-DNA.



#1Armenian: R1b1a2a-L23: R1b-M343+, M269 parallels+, L23+: xR1b1a2a1a2c-L21, R1b1a2a2a1-L943, R1b1a2a2c1. Rest of negatives are way downstream.

#2Armenian: CT: xG, a HIJK, I, J2a a L, E1a, E2, a E1b1a1, a E1b1b1a
J1a2a-Z1842+, xJ2a-M410
No calls in T
xQ1b, a Q1a, Q1a2a
H,C, R, K, N, O negatives are all downstream.

#3Armenian: E1b1b1b2a-CTS677:
E1b1b1b2a-CTS677 parallel to M123+.
xa E1b1b1b2a1. E1b1b1b2a1a-CTS877/L789/PF1512+ according to 10/13/2011 tree.
xR1, R1b, R1a1, R2

#4Armenian: R1b1-M415: R1, R1b, R1b1-M415+: R1b negatives are way downstream.

#5:Armenian: J:
HIJK+
xa R1a, a R1b1a2a2c
a J+. J negatives very downstream
xI, G, E, L, T, N, O
xQ1b, Q1a1.



https://docs.google.com/document/d/1Txjf7qEHsRUu9-o5KpKJvLAF465gr4HBekzcX__Y2hU/edit

[Faklon]

Yamna I2a?
Do we know the exact date and locations of Armenian samples?



Seems that J rampaged through the Middle East in the Iron Age.

J2b at least (http://cdn.eupedia.com/images/content/Haplogroup-J2b.gif), possibly.

[Vesuvian Sky]

I wonder if there are any new Gedmatch kits associated with these. Particularly Sintashta and some of the Anatolian samples.:icon_ask:

[curupira]

From the study:

http://i59.tinypic.com/2pys0nt.png
http://i58.tinypic.com/4iylox.png
http://i61.tinypic.com/1zdugif.png
http://i59.tinypic.com/111ujvq.jpg

[drgs]

"Populations in northern and central Europe were composed of a mixture of the earlier hunter-gatherer and Neolithic farmer groups, but received ‘Caucasian’ genetic input at the onset of the Bronze Age (Fig. 2). This coincides with the archaeologically well-defined expansion of the Yamnaya culture from the Pontic-Caspian steppe into Europe (Figs 1 and 2). This admixture event resulted in the formation of peoples of the Corded Ware and related cultures, as supported by negative ‘admixture’ f3 statistics when using Yamnaya as a source population (Extended Data Table 2, Supplementary Table 12). Although European Late Neolithic and Bronze Age cultures such as Corded Ware, Bell Beakers, Unetice, and the Scandinavian cultures are genetically very similar to each other (Fig. 2), they still display a cline of genetic affinity with Yamnaya, with highest levels in Corded Ware, lowest in Hungary, and central European Bell Beakers being intermediate (Fig. 2b and Extended Data Table 1)."

“Now we see the Yamnaya is not only spreading north into Europe; they’re also spreading east, crossing the Urals, getting all the way into central Asia, all the way into the Altai, between Mongolia, China, and Siberia,” says evolutionary biologist Eske Willerslev of the University of Copenhagen..."

"We find that the Bronze Age in Asia is equally dynamic and characterized by large-scale migrations and population replacements. The Early Bronze Age Afanasievo culture in the Altai-Sayan region is genetically indistinguishable from Yamnaya, confirming an eastward expansion across the steppe (Figs 1 and 3b; Extended Data Fig. 2b and Extended Data Table 1), in addition to the westward expansion into Europe. Thus, the Yamnaya migrations resulted in gene flow across vast distances, essentially connecting Altai in Siberia with Scandinavia in the Early Bronze Age (Fig. 1)."

"Interestingly, the Bronze Age steppe cultures showed the highest derived allele frequency among ancient groups, in particular the Yamnaya (Extended Data Fig. 7), indicating a possible steppe origin of lactase tolerance."

"From the beginning of 2000 BC, a new class of master artisans known as the Sintashta culture emerged in the Urals, building chariots, breeding and training horses (Fig. 1), and producing sophisticated new weapons18. These innovations quickly spread across Europe and into Asia where they appeared to give rise to the Andronovo culture19,20 (Fig.1)."

[Austrvegr]

So the truest living Aryans are Russians after all.


http://s22.postimg.org/64lijqhu9/pca12.png

http://s18.postimg.org/lsd2lytah/image.jpg




http://i1155.photobucket.com/albums/p546/meon_/aR1ans1.jpg

[Artek]

Staff of Yfull.com will take care of .BAM files to mine as many y-SNPs as possible.

[Rochefaton]

Here (http://www.filedropper.com/ancientarmeniansintashtak12boraclev1) is an OpenOffice-based spreadsheet that acts as the basic Oracle function from Gedmatch. It will currently let you compare your Dodecad K12b results to those of the MBA Armenian samples' average, the LBA Armenian samples' average and to the results of one Sintashta sample (kit# M690970). I will update the spreadsheet as more samples become available.

Here are my current results:

Armenia MBA: 29.7703461522
Armenia LBA: 39.4656917109
Sintashta M690970: 17.0491420628

I'm obviously much closer to the Sintashta sample than I am to either Armenian group, but my values are also 25% closer to the MBA Armenians than to the LBA samples.

[ALL]

Quite a distance R1b-CTS-7763 in B.A Armenia and recently found in Bhutan.


From the study:

http://i59.tinypic.com/2pys0nt.png
http://i58.tinypic.com/4iylox.png
http://i61.tinypic.com/1zdugif.png
http://i59.tinypic.com/111ujvq.jpg
http://www.nature.com/nature/journal/v522/n7555/full/nature14507.html
SAMEA3325367 RISE397 Kapan LBA Armenia Male 1048 BC 855 BC R1b1a2a2
]RISE397_Kapan_LBA_Armenia R1b>M269>L23>Z2103>Z2106>CTS7763 is a good catch by smal. R1b-CTS7763 is very specific branch of the R1b-Z2103 haplogroup. Most of us a year or two ago would have guessed Armemian R1b to be R1b-L584 or maybe R1b-L277.
The R1b-M269 (P312- U106-) DNA Project (aka ht35 Project) Phylogenetic Tree (https://www.familytreedna.com/groups/ht-3-5new/about/results) that smal has built has been very useful with these ancient samples.
[I]SAMEA3325367 RISE397 Kapan LBA Armenia Male 1048 BC 855 BC

]RISE397_Kapan_LBA_Armenia R1b>M269>L23>Z2103>Z2106>CTS7763 Male1048 BC 855 BC R1b1a2a2

http://oi58.tinypic.com/vgopvq.jpg

1KGP-NA18645 (Han Chinese in Beijing, China)
H2014-bhu-1953 (Bhutanese)
http://www.kumbarov.com/ht35/R1b1a2_ht35_project_tree_16_05_25_2015.pdf

[Artek]

1. Sintashta, Bulanovo, Russia, 2298 BC - 2045 BC, R1a-Z645>Z93>Z94>Z2124 !!
2. Sintashta, Stepnoe VII, Russia, 2126 BC - 1896 BC, R1a-Z645>Z93(?)
3. Andronovo, Kytmanovo,Russia, 1446 BC - 1298 BC, R1a-Z645>Z93(?)
4. Karasuk, Sabinka II, Russia, 1416 BC - 1268 BC, R1a1a1 (M417) Z93(?)
5. Meshovskaya, Kapova cave, Russia, nd, R1a1a1 (xZ283) Z93?
6. Late Bronze Age, Afontova Gora, 926 BC - 715 BC, R1a1a1 (M417) Z93(?)
7. Corded Ware, Tiefbrunn, Germany, 2880 BC - 2630 BC, R1a1a1(M417)
8. Corded Ware, Tiefbrunn, Germany, 2868 BC - 2580 BC, R1a1a1 (M417)
9. Corded Ware, Bergrheinfeldt, Germany, 2829 BC - 2465 BC, R1a1a1 (xZ647) CTS4385(?)
10. Corded Ware, Łęki Małe, Poland, 2286 BC - 2048 BC, R1a1a1 (M417)
11. Corded Ware/Battle-Axe, Viby, Sweden, 2621 BC - 2472 BC, R1a1a1 (M417)
12. Nordic Late Neolithic, Marbjerg, Denmark, 2191 BC - 1972 BC, R1a1a1 (M417)
13. Nordic Late Neolithic, Kyndel, Denmark, 2651 BC - 2292 BC(?), R1a1a1 (M417)
14. Late Bronze Age, Turlojiske, Lithuania, 908 BC -485 BC, R1a1a1 (M417)
15. Iron Age, Sabinka 2, Russia, 396 BC - 209 BC, R1a1a1 (xZ283) Z93(?)

To be updated in few days, hopefully.

[Graham]

So Sintashta R1a Z2124. Which is more of a Eurasian Y-DNA in todays world. But the Autosomals are looking very European.

[Graham]

Sintashta looks Northern Europe with the ANE link..


Sintashta Oracles
K13 Eurogenes
88.2% Swedish + 11.8% Tabassaran = 9.56
K15 Eurogenes
88.5% West_Norwegian + 11.5% Tabassaran = 6.76
Dodecad K12b
88.2% Swedish + 11.8% Balochi = 5.33


Dodecad K12b

Gedrosia 16.61%
Siberian 0.42%
Northwest_African 0.09%
Southeast_Asian -
Atlantic_Med 26.36%
North_European 53.75%
South_Asian -
East_African -
Southwest_Asian -
East_Asian -
Caucasus 1.86%
Sub_Saharan 0.90%

Chariot Spread
https://upload.wikimedia.org/wikipedia/commons/7/77/Chariot_spread.png
R1a Spread
http://cache.eupedia.com/images/content/R1a_migration_map.jpg

[noricum]

Sintashta M690970 eye colour

http://i63.photobucket.com/albums/h149/Christian_Unterberger/sintashta_zpsmgzpzoro.jpg (http://s63.photobucket.com/user/Christian_Unterberger/media/sintashta_zpsmgzpzoro.jpg.html)

[Artek]

Sintashta M690970 eye colour

http://i63.photobucket.com/albums/h149/Christian_Unterberger/sintashta_zpsmgzpzoro.jpg (http://s63.photobucket.com/user/Christian_Unterberger/media/sintashta_zpsmgzpzoro.jpg.html)
http://i938.photobucket.com/albums/ad228/CeltHussar/My%20eyes%20prediction_zpsn6dvy2kw.jpg

Basis is similar, some traits are different. Predictor works very accurate for me, I wonder if the same is for Sintashta sample. Depends on what wasn't found due to the lower coverage.

[curupira]

I'd even say a R1b-Centum vs a R1a-Satem division suddenly becomes reasonable again. People already foresaw it long ago but now it seems it could be true indeed.

This is what the poster DMXX said on Anthrogenica.


A wayward thought just occurred to me.

The Yamnaya and Afanasievo autosomal results are practically identical to one another. The former has been shown to be predominantly Y-DNA R1b consistently (Haak et al., this study) with the latter having rumours of being Y-DNA R1b. Relative to the other Pontic-Caspian steppe cultures sampled, these two appear to be the oldest.

Conversely, the Corded Ware, Sintashta and Andronovo autosomal results are also exceedingly similar to one another. Y-DNA R1a appears to be the common uniparental signal across all three. The latter two are consistently designated as "late" IE cultures (Sintashta and Poltkava immediately preceding Andronovo).

This is a clear over-simplification of the ethnogenesis which must have taken place on the steppes (particularly given the time and space differentiation between CW and Sintashta, alongside the numerous derived subcultures typically ascribed to specific proto-languages potentially having their own individual development)...

But, are we now in a position to generally posit:
1) Y-DNA R1b and a Yamnaya-Afanasievo autosomal profile typifying the early stages of PIE,
2) Y-DNA R1a and a CW-Sintashta autosomal profile representing the later stage(s) of PIE?

This happens to be in accordance with Anthony & Ringe's recent IE phylogeny (http://2.bp.blogspot.com/-Et6SS_bTFtc/VM740osEIeI/AAAAAAAAJ5I/3JmZBNHvNFU/s1600/annurev-linguist-030514-124812.f2.jpeg).

(P.S. To my distant R1 cousins, check the profile <----- R2-D4 (https://farm3.staticflickr.com/2392/2331188256_3229742c5d_o.jpg) has no personal stake in this R1valry :) )
http://www.anthrogenica.com/showthread.php?4640-Population-genomics-of-Bronze-Age-Eurasia-(Allentoft-et-al-2015)&p=89651&posted=1#post89651

[Hevo]

First R1b U106* has been found in Swedish Battle Axe Culture.:cool: (a Corded Ware subgroup)

https://drive.google.com/file/d/0B9vzsK0Ig1mNdlB4LTgzN2ZMNW8/view

[Nurzat]

we J2a's conquered Altai :odinsleipnir2: Uyghurs are still 10% J2a1b*

[Graham]

K15

Swedish Battle Axe Culture

1 North_Sea 39.93
2 Atlantic 18.64
3 Eastern_Euro 16.41
4 Baltic 15.82
5 West_Med 7.64
6 South_Asian 1.07
7 Sub-Saharan 0.36
8 Oceanian 0.12

Single Population Sharing:

# Population (source) Distance
1 North_Swedish 5.69
2 Swedish 6.66
3 Norwegian 6.81
4 West_Norwegian 8.26

80.4% West_Norwegian + 19.6% Erzya = 5.15
77.2% North_Swedish + 22.8% West_Norwegian = 5.38
69.4% North_Swedish + 30.6% Norwegian = 5.38

[Graham]

RISE174, Sweden Iron Age, 427-611 AD

F999943

We have a Swede that is only a few hundred years prior to the Viking Age. Have fun.

[Longbowman]

RISE174, Sweden Iron Age, 427-611 AD

F999943

We have a Swede that is only a few hundred years prior to the Viking Age. Have fun.

Apparently he's E-Z14919 and W1.

[Jackson]

RISE174, Sweden Iron Age, 427-611 AD

F999943

We have a Swede that is only a few hundred years prior to the Viking Age. Have fun.

84k SNPs so fairly good.

Looks like they were just more northern than a modern Swede, but still close to the modern population unsurprisingly:
K13
1 North_Atlantic 55.26
2 Baltic 35.18
3 West_Med 5.1
4 West_Asian 3.55
5 Amerindian 0.91

1 Swedish 8.21
2 Norwegian 8.46
3 North_Swedish 10.34
4 Danish 10.84
5 North_Dutch 10.92
6 North_German 12.03
7 Irish 12.52
8 Orcadian 12.54
9 West_Scottish 12.76
10 Southeast_English 14.61

1 86.2% Norwegian + 13.8% La_Brana-1 @ 7.77
2 68.1% Norwegian + 31.9% North_Swedish @ 7.85
3 87.9% Norwegian + 12.1% Finnish @ 7.95
4 88.8% Norwegian + 11.2% Estonian @ 7.96
5 84.3% Norwegian + 15.7% Southwest_Finnish @ 7.96

[Graham]

I dont know what E-Z14919 is in todays world? & how much SNPs are a good amount Jackson?

[Hevo]

RISE174, Sweden Iron Age, 427-611 AD

F999943

We have a Swede that is only a few hundred years prior to the Viking Age. Have fun.

Is there a list of the Gedmatch kits of these gnomes? They are all over the place.:/

[Graham]

Is there a list of the Gedmatch kits of these gnomes? They are all over the place.:/

Unfortunately everything inst together. But if you go into the Archaic DNA matches you can find some numbers and match them here. http://umap.openstreetmap.fr/en/map/ancient-human-dna_41837#4/49.87/93.78

And some more here.
https://www.dropbox.com/sh/mqp4gxtzpng3kom/AAC8ctJnRh-T_T73s81E7T75a/Samples.xlsx?dl=0

Plus some more out there somwhere.


Alberstedt I0118 - M467300
Halberstadt LBA I0099 M344778
Corded Ware I0104 M622615
Els Trocs Spain EN R1b1 I0410 M684242
Karelian-HG (EHG_Oleniy_Ostrov I0061) M652848
Yamna 1 (Yamnaya_Sok_River I0443) M951285
Yamna 2 (Yamnaya I0231) M020637
LBK I0054 - kit M206308
Corded Ware I0103 - kit M966366
Unetice I0047 - kit M370010
Bell Beaker I0112 M117132
Esperstedt I0172 M303249

[Not a Cop]

84k SNPs so fairly good.

Looks like they were just more northern than a modern Swede, but still close to the modern population unsurprisingly:
K13
1 North_Atlantic 55.26
2 Baltic 35.18
3 West_Med 5.1
4 West_Asian 3.55
5 Amerindian 0.91

1 Swedish 8.21
2 Norwegian 8.46
3 North_Swedish 10.34
4 Danish 10.84
5 North_Dutch 10.92
6 North_German 12.03
7 Irish 12.52
8 Orcadian 12.54
9 West_Scottish 12.76
10 Southeast_English 14.61

1 86.2% Norwegian + 13.8% La_Brana-1 @ 7.77
2 68.1% Norwegian + 31.9% North_Swedish @ 7.85
3 87.9% Norwegian + 12.1% Finnish @ 7.95
4 88.8% Norwegian + 11.2% Estonian @ 7.96
5 84.3% Norwegian + 15.7% Southwest_Finnish @ 7.96



Dude scores 74.2% WHG in ANE K7, so same as ancient Brits ancient Swede is a bit more European compared to modern ones.

[Jackson]

Dude scores 74.2% WHG in ANE K7, so same as ancient Brits ancient Swede is a bit more European compared to modern ones.

Makes sense, as since those periods both areas (although Britain more than Sweden i think) have had increasing influence from the Mediterranean world, or at least from the south/mainland Europe.

[Graham]

How does Hinxton 4 Iron Age Celtic Briton compared to the pre Viking Era Swede. Noise omitted.



British Iron Age K7
WHG-UHG 65.59%
ANE 16.62%
ENF 14.22%
ASE 1.52%

Eurogenes K13
1 Irish 2.72
2 West_Scottish 2.78
Eurogenes K15
1 Irish 4.3
2 West_Scottish 5.35
MDLP K23b
1 Irish 4.47
2 English 4.71


Swedish Iron Age K7
WHG-UHG 74.20%
ANE 17.66%
ENF 5.92%
ASE 1.91%

Eurogenes K13
1 Swedish 8.21
2 Norwegian 8.46
Eurogenes K15
1 Danish 8.91
2 North_Dutch 9.13
MDLP K23b
1 Norwegian_West 3.09
2 Icelandic 3.10

[Artek]

RISE494. Karasuk. Z93>Z94>Z2124>Z2125>Z2123>Y934>YP520+
RISE392. Stepnoe VII. Sintashta, Z93>Z94>Z2124>Z2125>Z2123>Y877+ Y939-
RISE512. Kytmanovo. Andronovo Z93>Z94>Z2124 (Z2121+, level Z2124) Y877-
RISE386. Bulanovo. Sintashta Z93>Z94>Z2124>Z2125>S23592>YP1456 (YP1460+, level YP1456)
Kapova cave.
RISE525. Meshovskaya. Z283(-? false negative)>Z282>Z280>CTS1211>YP343>YP340>YP371>Y11162 (Y11175+, level Y11162), Y11171- (Slavic skeleton?)

RISE446 (Bergrheinfeld/Corded Ware/Germany) M417>CTS4385>L664>S3477 (S3479+ S3490- YP233-)
RISE61 (Kyndel/Nordic LN/Denmark) 2851 BC - 2492 BC Z282>Z284>Z287>CTS8401>Z281 (Z281+ S6752- S3238- S3225-)