Ancient genomes from present-day France unveil 7,000 years of its demographic history

[dududud]

Link: https://cvws.icloud-content.com/B/AacoulFC9EelvPHiiGfIkHU9gmtyAcVCoZM7BSEjxAMT-Fl-FoklpMdq/10.1073%40pnas.1918034117.pdf?o=Ag-Se1c8gb7zI_zyoZuQtGDT7IXFhhXtl5W0n7wcFx38&v=1&x=3&a=CAogpjDurrikKBTo9KjqLq9K45mxLZ4-b0xF-42DsUde2PsSbxCmzu2SpS4YxsWkk6UuIgEAUgQ9gmtyWgQlpMd qaifzzyiAE7qwXEOBuc3wYyL8j2NSbsCQ9ZeKkm7jZp1YAtO3w XmU9JlyJ4-OeZkegxtYcPirPBgx_V1bzBqxT3OBeMXp_znCF-5nCavHkWIA2A&e=1590520521&fl=&r=c3f278af-8b8e-40c2-868c-6c70be0d98c2-1&k=9uJUtdTyyK5dggAAjLWiAQ&ckc=com.apple.clouddocs&ckz=com.apple.CloudDocs&p=45&s=xI1MBKS227gemWYnb-Ar8HMgLgM&cd=i

https://image.noelshack.com/fichiers/2020/22/2/1590520133-3939nln4.jpg

[dududud]

Results and Discussion
All analyzed individuals belong to well-defined archaeological
contexts from the Mesolithic Period to the Iron Age and 40 were directly radiocarbon-dated for this study (Dataset S2). We built sequencing libraries for 243 individuals and enriched them for mitochondrial genomes and a panel of 120 selected SNPs across the nuclear genome (Dataset S1). Along with complete mitochondrial genomes of 223 individuals and Y-chromosome haplogroups of 62 male individuals, we report 58 ∼0.05 to 0.5× genomes sequenced from uracil-DNA glycosylase (UDG)-treated libraries. We combined our genotype data with either complete genomes or 1,240k genome-wide polymorphisms in ancient individuals and genotypes of modern individuals from Europe, the Caucasus, and the Near East genotyped on the Affymetrix Human Origins array (3, 5, 9, 10, 12, 13, 21, 22). Complete mitochondrial genomes were retrieved from this ancient dataset and were grouped based on temporal and geographic proximity, often translating into cultural classifications (SI Appendix, Fig. S3-1). Contamination estimates derived from X-chromosome polymorphisms in male individuals were consistently low (between 0.68 ± 1.26 and 3.55 ± 0.62% depending on the method considered), with only one individual displaying more than 5% X-chromosome contamination (PIR3037AB) (Dataset S1).


The French Mesolithic Substrate.
We generated genomic data for five Mesolithic individuals from Les Perrats cave (Agris, Charente) dated from 7177 to 7057 calibrated years BCE (calBCE) including low-coverage shotgun sequencing for three of them
(0.111 to 0.134×). All possess haplogroup U5b, demonstrating in the territory of present-day France the presence of a Mesolithic substrate akin to that described elsewhere, where it was characterized by haplogroups U5b (85%) and, to a lesser extent, U5a (15%) (3, 23). Mesolithic hunter-gatherers (HGs) discovered in France are located near the edge of a principal-component analysis (PCA) plot obtained from genome-wide genotypes, falling near the Mesolithic individuals from western Europe (5, 6, 9) (Fig. 1B and SI Appendix, Fig. S4-3). While the ancestry from an ∼14,000-y-old individual from Villabruna in Italy was found to be dominant in Holocene HGs across western and central Europe (3), recently published studies conducted on ancient Iberians highlighted the survival of Magdalenian-associated ancestry (11).
This dual ancestry is found in all Iberian HGs (ranging from 23.7 to 75.3%), while in most other regions only the Villabrunaassociated ancestry remained. In order to investigate the proportions of ancestry in French HGs derived from these two Late Pleistocene lineages, we used qpAdm to model it as a mixture of two sources: ∼15,000-y-old GoyetQ2 from Belgium, the leastadmixed individual associated with the Magdalenian complex in Europe sequenced so far (11), and Villabruna. We found that French HGs from Les Perrats harbored relatively high proportions of GoyetQ2 ancestry ranging from 31.3 to 45.6% (Fig. 1C), comparable to proportions described in the La Braña or Canes1 Mesolithic individuals from Spain, suggesting a late survival of Magdalenian-associated ancestry in HGs outside the Iberian Peninsula.


Successive Waves of Migration and Admixture over the Course of the Neolithic in France.

The arrival of an Anatolian Neolithicassociated ancestry component at the onset of the Neolithic is clearly visible in our dataset from both uniparentally inherited markers and at the genome-wide level. Maternal lineages from Neolithic farmers in France (from ∼5300 BCE onward) are more diverse than those from HGs, and display variable frequencies across the time transect (SI Appendix, Fig. S3-2). Early and Middle Neolithic individuals share more affinity with present-day
southern Europeans (SI Appendix, Fig. S4-2), and their genetic variation is encompassed within that of contemporaneous European populations (Fig. 1B). Three of our individuals associated with the LBK culture, which stems from the Danubian Neolithic current, cluster genetically with other central European Early Neolithic (EN) individuals and share drift with other early farmers associated with the LBK cultural complex (SI Appendix, Fig. S4-6). Values of D (Mbuti, France_EN; western huntergatherers [WHGs], Anatolia_Neolithic) are consistent with those observed in LBK-associated individuals, with the exception of Mor6 (∼7,100 calB.P.) in northeastern France (Fig. 2B).Despite a consistent dating and cultural assignment to the LBK, this individual falls within the genetic diversity of Iberian_EN individuals and harbors the highest proportion of shared alleles with WHGs. After testing the potential origin of this ancestry with qpAdm, we found the best fit to be with a two-source ancestry model between Anatolia_Neolithic and GoyetQ2 (P =0.0911155), a situation so far only described in Early Neolithic individuals from the Iberian Peninsula. This is the northern-most description of such ancestry. To date, the distribution of the Magdalenian-associated ancestry in the HG population in western Europe that admixed with the Neolithic farmers in Europe is not clear because of the scarcity of genetic data from Mesolithic HGs. The observation of this ancestry in the Iberian Peninsula has been interpreted as a feature distinguishing the Cardial from the LBK Neolithic migrants (12).
The observation that the ancestry of GoyetQ2 was found alongside Villabruna ancestry in HGs from Les Perrats in western France (∼9,100 calB.P.), as well as in HGs from Rigney1 (central-eastern France; ∼15,500 calB.P.) and from two caves in
southwestern Germany, Hohlefels (∼15,000 calB.P.) and Burkhardtshöhle (∼14,600 calB.P.), rather indicates that this mixed
ancestry is characteristic of western European HGs (11). Thus, the admixture between HGs with GoyetQ2 ancestry and Neolithic migrants is not necessarily characteristic of the Cardial Neolithic migration wave, as it seems to have occurred as well in more northern locations in western Europe. While the only male Mesolithic individual from our dataset
was assigned to Y-haplogroup I, LBK individuals from France showed relative diversity, belonging to Y-haplogroups C1a2,
G2a, and H2. G2a and H2 have been described in various European Early Neolithic contexts in both central Europe and
Iberia (3, 6, 24). I2 haplogroups are likely to have been introduced into the Neolithic pool through admixture with huntergatherers.

In eastern France, the transition into the Middle Neolithic is represented by individuals from the Grossgartach culture, that derives from the Danubian sphere (∼4700 to 4500 BCE; SI Appendix, Fig. S1-1). Their proximity to central European contemporaneous individuals on PCA plots generated from both mitochondrial haplogroup frequencies and genome-wide markers suggests genetic homogeneity within the Danubian current (Fig. 1B and SI Appendix, Fig. S3-3). Interestingly, the BUCH2 individual harbors mixed GoyetQ2 and Villabruna HG ancestry (Fig. 1C), suggesting again that the GoyetQ2 ancestry is not a specific signature of southwestern Europe. However, we can observe a subtle shift away from LBK individuals and toward WHGs, reflecting an increase in shared alleles with the latter. This echoes observations from other ancient populations across Europe (Figs. 1C and 2A), yet with a higher proportion. This phenomenon seems to appear in France very early and becomes stronger as the Neolithic progresses, since individuals from French Middle Neolithic sites share a higher proportion of ancestry with HGs (Fig. 2A) and display Mesolithic signature haplogroups (e.g., U5b) at variable frequencies across the territory (SI Appendix, Fig. S3-2). Predominant in the Champagne region of northeastern France, these haplogroups suggest variable proportions of admixture between autochthonous hunter-gatherers and farmers. During the second half of the Middle Neolithic (MN2, ∼4500 BCE), individuals from northern and southern France are poorly distinguished on the PCA plot (Fig. 1B). Unlike in Iberia, the migration wave from which the Neolithic ancestry in ancient populations from France originated could not clearly be established using D statistics comparing the ancestry proportion between Iberia_EN and LBK_EN, as most individuals display nonsignificant values (SI Appendix, Fig. S4-6) (12). Using qpAdm, we investigated the ancestral source of the HG ancestry in individuals from French MN2 sites. Only individuals from southern France harbor varying levels of GoyetQ2-like ancestry, suggesting admixture with different populations of hunter-gatherers (Fig. 1C).

[XenophobicPrussian]

Told you mothercluckers, not a single one of the southern shifted IA Gauls was actually from the south. :laugh: They're all from Alsace and Paris area, including the Iberian clustering ones.

Rhaetians? :laugh:

muh genetic continuity

[Aren]

Holy, a fully Yamnaya scoring individual from Bronze Age France(!). And he's R1b, anyone know his final subclade? The link doesn't work btw anymore.

[dududud]

Told you petit coquin, not a single one of the southern shifted IA Gauls was actually from the south. :laugh: They're all from Alsace and Paris area, including the Iberian clustering ones.

Rhaetians? :laugh:

muh genetic continuity


There are samples from the South. You seem blind and stupid.

[Lucas]

Only two IA are from Hauts de France
Probably only this (Att226) is in G25.

[Lucas]

The link doesn't work btw anymore.

https://sci-hub.ren/https://doi.org/10.1073/pnas.1918034117

[Lucas]

Suplemental https://www.pnas.org/content/pnas/suppl/2020/05/20/1918034117.DCSupplemental/pnas.1918034117.sapp.pdf

https://i.imgur.com/8s5oTZN.png



Bronze and Iron Age France share a common space in thePCA plot, both shifted toward modern central Europe and falling within the genetic diversity of Bronze Age Britain and central Europe (Fig. 1B), with a homogenization of the steppe component (Figs. 1Cand 2C). In contrast to what was described for central Europe (9), there is no further shift toward eastern Eurasian genotypes during the Iron Age (9). Instead, the steppe component, heterogeneously distributed between individuals during the Bronze Age (ranging from 30 to 70%), becomes homogeneous (Figs. 1Cand 2C), and individuals from the Hallstatt and La Tène culture in the French territory display similar affinities toward both modern and ancient populations. This could indicate that the transition from the Bronze Age to the Iron Age in France was mostly driven by cultural diffusion, without major gene flow from an external population. This would be consistent with an archeological and linguistic hypothesis proposing that the Celts from the second Iron Age descended from populations already established in western Europe, within the boundaries of the Bell Beaker cultural complex (26–29). It is important to mention, however, that due to the relative genetic homogeneity among European populations by the Bronze Age, subsequent migrations between different parts of Europe could easily remain unnoticed at this level of coverage.

[XenophobicPrussian]

There are samples from the South. You seem blind and stupid.
Don't bother me with your English second language or coping mechanisms right now Frenchie(or should I say Germanic+Celt+Roman+native Beaker).

COL153i, ERS88, Jeb8, NOR2B6, NOR3-15, NOR4, all the southern shifted(SW/S French or Iberian clustering) samples that were out already, are all in Alsace or Paris area France, not a single one in southern France, according to the image.

Game over.

[Lucas]

OK:) New averages comparing to modern pops.

Target: FRA_IA_north (no big difference than today)
Distance: 2.7289% / 0.02728949 | ADC: 1x
86.2 Belgian
8.4 French_Paris
5.4 French_Brittany

Target: FRA_IA_central (southern shifted central French)
Distance: 1.3569% / 0.01356942 | ADC: 1x
63.2 French_Occitanie
36.8 French_Nord

Target: FRA_IA_south YES
Distance: 2.8076% / 0.02807648 | ADC: 1x
68.2 French_Brittany
24.2 Orcadian
7.6 French_Seine-Maritime





north

FRA_IA:ATT26,0.1161,0.132019,0.044877,0.039083,0.0 38776,0.013108,-0.006815,0.011999,0.014112,0.020957,-0.001299,0.007793,-0.024678,-0.011836,0.007872,0.007027,0.006128,0.008361,0.010 307,-0.013632,0.003993,0.00779,0.020706,0.000482,0.0087 42
FRA_IA:BFM265,0.122929,0.148267,0.05242,0.037468,0 .047086,0.018128,-0.00235,-0.000231,0.008795,0.012392,-0.007957,0.005695,-0.015312,-0.007844,0.020765,-0.003182,-0.016428,0.009122,0.006034,0.007504,-0.002246,0.000124,0.001109,-0.00012,0.002036
average,0.1195145,0.140143,0.0486485,0.0382755,0.0 42931,0.015618,-0.0045825,0.005884,0.0114535,0.0166745,-0.004628,0.006744,-0.019995,-0.00984,0.0143185,0.0019225,-0.00515,0.0087415,0.0081705,-0.003064,0.0008735,0.003957,0.0109075,0.000181,0.0 05389


central
FRA_IA:COL11,0.122929,0.13405,0.058454,0.064923,0. 037238,0.020917,-0.008225,0.01223,0.008795,0.011116,-0.002273,0.002548,-0.016353,-0.003991,0.017508,-0.008221,-0.014603,-0.005954,0.006662,0.005378,0.006489,0.008161,0.010 106,-0.001446,0.000599
FRA_IA:COL153A,0.130897,0.158423,0.045254,0.02584, 0.036622,0.012271,0.006815,0.008538,0.003272,0.015 126,0.009419,-0.007194,0.001189,-0.000963,0.005022,0.003182,-0.013038,0.00038,0.016592,0.001,-0.015847,0.007543,0.000863,-0.018798,0.005389
FRA_IA:COL153i,0.120652,0.127957,0.046763,0.024548 ,0.045239,-0.006136,0.015276,0.004154,0.015748,0.034078,-0.00341,0.017235,-0.022448,-0.010459,0.012622,-0.026518,0.005476,0.008868,-0.001131,0.015883,-0.000749,-0.000742,-0.010476,0.003253,0.008143
FRA_IA:ERS86,0.122929,0.161469,0.040729,0.030039,0 .040623,0.008646,-0.010105,0.005077,0.006954,0.007289,-0.00065,0.005845,-0.010852,-0.002752,0.005157,0.021745,-0.004172,0.009755,0.011313,-0.005753,0.000624,0.012365,-0.004683,-0.012893,0.00012
FRA_IA:ERS88,0.130897,0.155376,0.044123,0.005168,0 .033237,0.000837,-0.004935,-0.005769,0.011249,0.02442,-0.004872,-0.001499,-0.023042,-0.016239,0.012079,-0.017104,-0.014603,-0.006461,-0.014204,-0.002251,-0.002496,0.000742,0.001356,0,0.000239
FRA_IA:ERS1164,0.124067,0.121864,0.052797,0.030039 ,0.033237,0.010877,0.011516,0.00923,0,0.019135,-0.002598,-0.01154,-0.01115,-0.012799,0.009908,0.006364,-0.002738,0.00038,0.001257,-0.004627,-0.007487,0.003957,0.002218,-0.011809,0.005389
FRA_IA:Jeb8,0.118376,0.138112,0.05506,0.027132,0.0 45239,-0.008367,-0.00329,0.003461,0.012271,0.010023,-0.00341,0.007194,-0.020812,-0.010046,0.016694,0.007425,-0.000391,-0.001647,0.000377,0.005378,-0.003868,-0.00136,-0.009983,-0.003615,0.007065
FRA_IA:NOR2B6,0.120652,0.147252,0.044877,0.005814, 0.0397,0.003347,-0.010105,0.007384,0.016975,0.014943,0.010718,0.011 09,-0.008771,-0.004542,0.012215,0.018165,0.027772,0.004941,0.005 405,0.002876,0.006364,-0.007543,-0.001109,-0.00253,-0.007784
FRA_IA:NOR3-6,0.133173,0.146236,0.050157,0.046189,0.029852,0.0 17291,-0.007755,0.005077,0.012885,0.001276,-0.002598,0.015137,-0.022745,-0.013212,0.020087,-0.007823,-0.021383,0.007095,-0.001006,0.006753,0.004866,0.005193,-0.002835,-0.007953,-0.000239
FRA_IA:NOR3-15,0.117238,0.149283,0.061471,0.022287,0.043393,-0.005299,0.005405,-0.001615,0.013703,0.018224,-0.005196,0.00045,-0.014569,-0.012111,0.022394,-0.005171,-0.019558,-0.006841,-0.004902,0.013381,-0.002121,-0.002844,-0.015652,-0.003615,0.002395
FRA_IA:NOR4,0.12862,0.151314,0.058454,0.001615,0.0 46778,-0.006136,-0.000705,0.014538,0.023929,0.031345,-0.013153,0.006594,-0.022448,-0.016377,-0.016965,0.004773,0.005737,-0.010135,0.011941,0.005503,0.006239,0.006183,-0.011462,0.007471,-0.002994
average,0.1245845455,0.1446669091,0.0507399091,0.0 257812727,0.0391961818,0.0043861818,-0.0005552727,0.0056640909,0.0114346364,0.016997727 3,-0.0016384545,0.0041690909,-0.0156364545,-0.0094082727,0.010611,-0.0002893636,-0.0046819091,3.46363636363638E-005,0.0029367273,0.0039564545,-0.000726,0.0028777273,-0.003787,-0.0047213636,0.0016656364

south
FRA_IA:PEY53,0.133173,0.139128,0.050534,0.039083,0 .050163,0.010598,-0.006345,-0.000692,0.008181,0.005103,-0.001624,0.016935,-0.019326,-0.007844,0.029451,-0.001724,-0.008996,0.003674,0.005028,-0.007128,0.010606,0.005193,-0.01861,-0.006507,0.008023
FRA_IA:BES1248,0.12862,0.127957,0.064488,0.042636, 0.043085,0.027052,0.007755,-0.003461,0.001636,0.007836,0.010718,0.011839,-0.00892,-0.007569,0.016015,-0.000663,-0.001434,0.004687,-0.015461,0.004002,0.01984,-0.002597,-0.009367,0.008555,-0.016765
average,0.1308965,0.1335425,0.057511,0.0408595,0.0 46624,0.018825,0.000705,-0.0020765,0.0049085,0.0064695,0.004547,0.014387,-0.014123,-0.0077065,0.022733,-0.0011935,-0.005215,0.0041805,-0.0052165,-0.001563,0.015223,0.001298,-0.0139885,0.001024,-0.004371

[XenophobicPrussian]

Can someone try to find the dating on BES1248? He's the Y-DNA I1 IA sample clustering northern autosomally, yet is from coastal southern France.. I know all the IA samples are from 800-100 BC, but can't find specific for each sample.

Seems to me like he'd either be a Cimbri/Teuton(but this heavily depends on the dating) or I1 is from Bell Beakers and his wasn't the Germanic clade.

Btw, in their section on their qpADM models,
None of our Iron Age samples shows evidence of Greek or Roman ancestry in our dataset, if anyone wants to claim the south shifted samples are Italics rather than Hallstatt Celts, and Rome only held parts of southern France in 100 BC anyway.

[Lucas]

Can someone try to find the dating on BES1248? He's the Y-DNA I1 IA sample clustering northern autosomally, yet is from coastal southern France.. I know all the IA samples are from 800-100 BC.

Seems to me like he'd either be a Cimbri/Teuton(but this heavily depends on the dating) or I1 is from Bell Beakers and his wasn't the Germanic clade.

Btw, in their section on their qpADM models, , if anyone wants to claim the south shifted samples are Italics rather than Hallstatt Celts, and Rome only held parts of southern France in 100 BC anyway.

Anyway those "northern southerners" had to be displaced or diluted later.
Alsace got Germanic influx.
But Hauts de France stays the same since IA... If we can conclude using so small subset.

[Samnium]

OK:) New averages comparing to modern pops.

Difficult to conclude something with so little samples in the south and the north I would say, we have to know if these two samples in the south are "immigrants".

[J. Ketch]

OK:) New averages comparing to modern pops.

Target: FRA_IA_north (no big difference than today)
Distance: 2.7289% / 0.02728949 | ADC: 1x
86.2 Belgian
8.4 French_Paris
5.4 French_Brittany

Target: FRA_IA_central (southern shifted central French)
Distance: 1.3569% / 0.01356942 | ADC: 1x
63.2 French_Occitanie
36.8 French_Nord

Target: FRA_IA_south YES
Distance: 2.8076% / 0.02807648 | ADC: 1x
68.2 French_Brittany
24.2 Orcadian
7.6 French_Seine-Maritime


That's a big WTF, but it kind of makes sense. If true it may explain why Iberia had such a Northern shift in the Iron Age despite 'Celts' allegedly being SW Euro like. The alternative would have been that Celts almost totally replaced Bronze Age Iberians, which obviously isn't true.

[Lucas]

Difficult to conclude something with so little samples in the south and the north I would say, we have to know if these two samples in the south are "immigrants".

There are more samples which weren't added to G25. Someone can compare now in other calcs if those not added are also "northern" in the south...

[Lucas]

Can someone try to find the dating on BES1248? He's the Y-DNA I1 IA sample clustering northern autosomally, yet is from coastal southern France.. I know all the IA samples are from 800-100 BC, but can't find specific for each sample.

Seems to me like he'd either be a Cimbri/Teuton(but this heavily depends on the dating) or I1 is from Bell Beakers and his wasn't the Germanic clade.

Btw, in their section on their qpADM models, , if anyone wants to claim the south shifted samples are Italics rather than Hallstatt Celts, and Rome only held parts of southern France in 100 BC anyway.

On Anthrogenica someone post


I received a response from the scientist in charge of the sample BES1248. It’s from a site very much in the south of France at Bessan, La Monediére from the second century B.C.

Unfortunately, she couldn’t provide any more information regarding Y chromosome SNP calls. Based on the quality of the sample and the approach they took this is all we have to work with at this point in time. A different approach taken in the future that targets the Y chromosome could possibly yield better coverage.

Based on the above information I’m leaning more heavily now on the sample not actually being positive for I-S10350. However, since BES1248 is genetically more similar to people from the north of France maybe he wasn’t actually born in the South of France in which case I-S10350 could still be a possibility?
If the Teutones and Ambrones could fight the Romans in 102 BC at the Battle of Aquae Sextiae all the way south in Aix-en-Provence, France maybe BES1248 is not so far displaced after all?

[Eline]

Thanks for sharing this paper!

https://i.postimg.cc/TYjRxDvZ/unnamed.jpg

[Eline]

So, does this mean than the Gauls evolved from the Bell Beaker in NorthEast France, but sill inside France?

https://i.postimg.cc/DZxcSKYq/Urnfield-Culture.jpg

[Eline]

So, does this mean than the Gauls evolved from the Bell Beaker in NorthEast France, but sill inside France?

https://i.postimg.cc/DZxcSKYq/Urnfield-Culture.jpg
And later the original homeland of the Gauls (inside France or Belgium) was overrun by the Franks???

[Eline]

The biggest irony is that the native language of France (French) is not from the Gauls and neither from the Franks but from Romance/Italic. :crazy:

[gixajo]

Haplogroup changing when yamnaya appeared is quite interesting . Did you noticed it?

[Eline]

Haplogroup changing when yamnaya appeared is quite interesting . Did you noticed it?Yes, R1b came with the Gauls!

https://i.postimg.cc/3wHDjJyg/R1b.jpg

[gixajo]

Are the kits of the individuals used in G25 ancient models available in gedmatch ?

I am not interested in those exactly, but in one individual who was found in a dolmen near the village of my mother, that appeared casually in one of my results.

[gixajo]

Yes, R1b came with the Gauls!

g]


I guess is a joke.:)

No, it´s very interesting, is something already known time ago, but here we can see it pretty well, how the paternal side of Indoeuropeans supplants the previous one, but not in the same proportion maternal line of previous population.

[celticdragongod]

Distance to: CDG_scaled
0.03577431 FRA_IA:COL11
0.03728353 FRA_IA:BFM265
0.04170613 FRA_IA:NOR3-6
0.05040868 FRA_IA:BES1248
0.05207900 FRA_IA:PEY53
0.05493145 FRA_IA:ERS1164
0.05515775 FRA_IA:Jeb8
0.05654879 FRA_IA:ATT26
0.06078658 FRA_IA:NOR3-15
0.06590770 FRA_IA:COL153A
0.06621788 FRA_IA:ERS86
0.07314464 FRA_IA:COL153i
0.07362566 FRA_IA:ERS88
0.07989075 FRA_IA:NOR2B6
0.08481511 FRA_IA:NOR4

Target: CDG_scaled
Distance: 2.9967% / 0.02996738 | ADC: 2x
64.8 FRA_IA:COL11
35.2 FRA_IA:BFM265

Target: CDG_scaled
Distance: 2.9039% / 0.02903928 | ADC: 1x
54.8 FRA_IA:COL11
42.4 FRA_IA:BFM265
2.8 FRA_IA:BES1248

Target: CDG_scaled
Distance: 2.7499% / 0.02749911 | ADC: 0.5x
47.2 FRA_IA:COL11
36.6 FRA_IA:BFM265
16.2 FRA_IA:BES1248

Target: CDG_scaled
Distance: 2.7363% / 0.02736321 | ADC: 0.25x
44.8 FRA_IA:COL11
34.0 FRA_IA:BFM265
19.4 FRA_IA:BES1248
1.4 FRA_IA:ERS1164
0.4 FRA_IA:NOR3-6

Target: CDG_scaled
Distance: 2.7321% / 0.02732052
44.0 FRA_IA:COL11
30.0 FRA_IA:BFM265
20.0 FRA_IA:BES1248
2.4 FRA_IA:ERS1164
2.0 FRA_IA:NOR3-6
1.6 FRA_IA:NOR4

[Coastal Elite]

Distance to: DacoCeltic_scaled
0.03423691 average_Central
0.03927802 average_North
0.04420762 FRA_IA:ERS1164
0.04646285 FRA_IA:BFM265
0.04700995 FRA_IA:Jeb8
0.04750392 FRA_IA:ATT26
0.04986210 average_South
0.05193589 FRA_IA:ERS86
0.05219918 FRA_IA:NOR3-6
0.05453842 FRA_IA:ERS88
0.05529729 FRA_IA:PEY53
0.05555213 FRA_IA:COL153A
0.05668221 FRA_IA:NOR2B6
0.05874818 FRA_IA:BES1248
0.05875382 FRA_IA:COL11
0.06063956 FRA_IA:NOR3-15
0.06211210 FRA_IA:COL153i
0.06375474 FRA_IA:NOR4


Target: DacoCeltic_scaled
Distance: 2.8987% / 0.02898707 | ADC: 0.25x
22.6 FRA_IA:ERS1164
16.8 FRA_IA:ATT26
13.0 FRA_IA:NOR3-6
11.8 FRA_IA:ERS88
11.6 FRA_IA:BES1248
11.0 FRA_IA:NOR2B6
9.8 FRA_IA:NOR4
3.4 FRA_IA:COL153A

Target: DacoCeltic_scaled
Distance: 3.1461% / 0.03146141 | ADC: 0.5x
68.8 average_Central
14.4 FRA_IA:ATT26
9.8 FRA_IA:ERS1164
6.6 FRA_IA:BES1248
0.4 FRA_IA:NOR2B6

[gixajo]

OK:) New averages comparing to modern pops.







north

FRA_IA:ATT26,0.1161,0.132019,0.044877,0.039083,0.0 38776,0.013108,-0.006815,0.011999,0.014112,0.020957,-0.001299,0.007793,-0.024678,-0.011836,0.007872,0.007027,0.006128,0.008361,0.010 307,-0.013632,0.003993,0.00779,0.020706,0.000482,0.0087 42
FRA_IA:BFM265,0.122929,0.148267,0.05242,0.037468,0 .047086,0.018128,-0.00235,-0.000231,0.008795,0.012392,-0.007957,0.005695,-0.015312,-0.007844,0.020765,-0.003182,-0.016428,0.009122,0.006034,0.007504,-0.002246,0.000124,0.001109,-0.00012,0.002036
average,0.1195145,0.140143,0.0486485,0.0382755,0.0 42931,0.015618,-0.0045825,0.005884,0.0114535,0.0166745,-0.004628,0.006744,-0.019995,-0.00984,0.0143185,0.0019225,-0.00515,0.0087415,0.0081705,-0.003064,0.0008735,0.003957,0.0109075,0.000181,0.0 05389


central
FRA_IA:COL11,0.122929,0.13405,0.058454,0.064923,0. 037238,0.020917,-0.008225,0.01223,0.008795,0.011116,-0.002273,0.002548,-0.016353,-0.003991,0.017508,-0.008221,-0.014603,-0.005954,0.006662,0.005378,0.006489,0.008161,0.010 106,-0.001446,0.000599
FRA_IA:COL153A,0.130897,0.158423,0.045254,0.02584, 0.036622,0.012271,0.006815,0.008538,0.003272,0.015 126,0.009419,-0.007194,0.001189,-0.000963,0.005022,0.003182,-0.013038,0.00038,0.016592,0.001,-0.015847,0.007543,0.000863,-0.018798,0.005389
FRA_IA:COL153i,0.120652,0.127957,0.046763,0.024548 ,0.045239,-0.006136,0.015276,0.004154,0.015748,0.034078,-0.00341,0.017235,-0.022448,-0.010459,0.012622,-0.026518,0.005476,0.008868,-0.001131,0.015883,-0.000749,-0.000742,-0.010476,0.003253,0.008143
FRA_IA:ERS86,0.122929,0.161469,0.040729,0.030039,0 .040623,0.008646,-0.010105,0.005077,0.006954,0.007289,-0.00065,0.005845,-0.010852,-0.002752,0.005157,0.021745,-0.004172,0.009755,0.011313,-0.005753,0.000624,0.012365,-0.004683,-0.012893,0.00012
FRA_IA:ERS88,0.130897,0.155376,0.044123,0.005168,0 .033237,0.000837,-0.004935,-0.005769,0.011249,0.02442,-0.004872,-0.001499,-0.023042,-0.016239,0.012079,-0.017104,-0.014603,-0.006461,-0.014204,-0.002251,-0.002496,0.000742,0.001356,0,0.000239
FRA_IA:ERS1164,0.124067,0.121864,0.052797,0.030039 ,0.033237,0.010877,0.011516,0.00923,0,0.019135,-0.002598,-0.01154,-0.01115,-0.012799,0.009908,0.006364,-0.002738,0.00038,0.001257,-0.004627,-0.007487,0.003957,0.002218,-0.011809,0.005389
FRA_IA:Jeb8,0.118376,0.138112,0.05506,0.027132,0.0 45239,-0.008367,-0.00329,0.003461,0.012271,0.010023,-0.00341,0.007194,-0.020812,-0.010046,0.016694,0.007425,-0.000391,-0.001647,0.000377,0.005378,-0.003868,-0.00136,-0.009983,-0.003615,0.007065
FRA_IA:NOR2B6,0.120652,0.147252,0.044877,0.005814, 0.0397,0.003347,-0.010105,0.007384,0.016975,0.014943,0.010718,0.011 09,-0.008771,-0.004542,0.012215,0.018165,0.027772,0.004941,0.005 405,0.002876,0.006364,-0.007543,-0.001109,-0.00253,-0.007784
FRA_IA:NOR3-6,0.133173,0.146236,0.050157,0.046189,0.029852,0.0 17291,-0.007755,0.005077,0.012885,0.001276,-0.002598,0.015137,-0.022745,-0.013212,0.020087,-0.007823,-0.021383,0.007095,-0.001006,0.006753,0.004866,0.005193,-0.002835,-0.007953,-0.000239
FRA_IA:NOR3-15,0.117238,0.149283,0.061471,0.022287,0.043393,-0.005299,0.005405,-0.001615,0.013703,0.018224,-0.005196,0.00045,-0.014569,-0.012111,0.022394,-0.005171,-0.019558,-0.006841,-0.004902,0.013381,-0.002121,-0.002844,-0.015652,-0.003615,0.002395
FRA_IA:NOR4,0.12862,0.151314,0.058454,0.001615,0.0 46778,-0.006136,-0.000705,0.014538,0.023929,0.031345,-0.013153,0.006594,-0.022448,-0.016377,-0.016965,0.004773,0.005737,-0.010135,0.011941,0.005503,0.006239,0.006183,-0.011462,0.007471,-0.002994
average,0.1245845455,0.1446669091,0.0507399091,0.0 257812727,0.0391961818,0.0043861818,-0.0005552727,0.0056640909,0.0114346364,0.016997727 3,-0.0016384545,0.0041690909,-0.0156364545,-0.0094082727,0.010611,-0.0002893636,-0.0046819091,3.46363636363638E-005,0.0029367273,0.0039564545,-0.000726,0.0028777273,-0.003787,-0.0047213636,0.0016656364

south
FRA_IA:PEY53,0.133173,0.139128,0.050534,0.039083,0 .050163,0.010598,-0.006345,-0.000692,0.008181,0.005103,-0.001624,0.016935,-0.019326,-0.007844,0.029451,-0.001724,-0.008996,0.003674,0.005028,-0.007128,0.010606,0.005193,-0.01861,-0.006507,0.008023
FRA_IA:BES1248,0.12862,0.127957,0.064488,0.042636, 0.043085,0.027052,0.007755,-0.003461,0.001636,0.007836,0.010718,0.011839,-0.00892,-0.007569,0.016015,-0.000663,-0.001434,0.004687,-0.015461,0.004002,0.01984,-0.002597,-0.009367,0.008555,-0.016765
average,0.1308965,0.1335425,0.057511,0.0408595,0.0 46624,0.018825,0.000705,-0.0020765,0.0049085,0.0064695,0.004547,0.014387,-0.014123,-0.0077065,0.022733,-0.0011935,-0.005215,0.0041805,-0.0052165,-0.001563,0.015223,0.001298,-0.0139885,0.001024,-0.004371




I don´t know why I get better distances with central samples and not with South ones.



Target: gixajo_scaled
Distance: 2.5204% / 0.02520371
30.0 FRA_IA:ERS88
27.8 FRA_IA:NOR2B6
16.6 FRA_IA:NOR3-15
15.8 FRA_IA:COL153i
9.8 FRA_IA:NOR4

Target: gixajo_scaled
Distance: 2.5210% / 0.02520966 | ADC: 0.25x
30.8 FRA_IA:ERS88
27.6 FRA_IA:NOR2B6
17.0 FRA_IA:NOR3-15
15.2 FRA_IA:COL153i
9.4 FRA_IA:NOR4


Distance to: gixajo_scaled
0.03773062 averageCentral
0.04212763 FRA_IA:ERS88
0.04416381 FRA_IA:NOR3-15
0.04483023 FRA_IA:Jeb8
0.05128186 FRA_IA:NOR2B6
0.05148714 FRA_IA:COL153i
0.05272917 FRA_IA:NOR4
0.05487212 averageNorth
0.05512703 FRA_IA:BFM265
0.05940530 FRA_IA:ERS86
0.06212623 FRA_IA:ERS1164
0.06278213 FRA_IA:COL153A
0.06312113 FRA_IA:PEY53
0.06443228 averageSouth
0.06567193 FRA_IA:ATT26
0.06938475 FRA_IA:NOR3-6
0.07652085 FRA_IA:BES1248
0.08207717 FRA_IA:COL11

[Grace O'Malley]

Haven't had a chance to read this paper yet but will asap. So L21 in Bronze Age France. First continental L21. That's nice. Clarifying my post that is outside ancient samples from Britain and Ireland. I hope more of these type of studies come out.

[gixajo]

Haven't had a chance to read this paper yet but will asap. So L21 in Bronze Age France. First continental L21. That's nice.

Anyway, I don´t know if the criteria for grouping samples in North/Center/South is the best.

[Peterski]

Don't bother me with your English second language or coping mechanisms right now Frenchie

How does our Prussian Xeno Warrior Prince feel about his beloved Nordic Viking I1-M253 haplogrup being possibly descended from the ancient Frenchies?: :laugh:

https://www.theapricity.com/forum/showthread.php?323831-I1-found-in-Neolithic-France&p=6719591&viewfull=1#post6719591


(...) Target: FRA_IA_south YES
Distance: 2.8076% / 0.02807648 | ADC: 1x
68.2 French_Brittany
24.2 Orcadian
7.6 French_Seine-Maritime

^^^
So, anyone still remembers the "Breton outlier" from Empuries?:

Distance to: Iberia_Northeast_Empuries1:I8206
0.03406911 French_Brittany
0.03535686 English
0.03539862 Welsh
0.03577452 Dutch
0.03719681 Norwegian
0.03722799 Irish
0.03732860 Scottish
0.03754347 French_Pas-de-Calais
0.03770647 English_Cornwall
0.03812733 Belgian
0.03880635 Orcadian
(...)

By the way, these two could also be Gauls from South France:

Distance to: Iberia_Northeast_Empuries1:I8214
0.03024663 French_Occitanie
0.03292765 French_Auvergne
0.03293229 French_South
(...)

Distance to: Iberia_Northeast_Empuries1:I8203
0.03155570 French_Occitanie
0.03188408 Swiss_French
0.03199066 French_Auvergne
(...)



south
FRA_IA:PEY53,0.133173,0.139128,0.050534,0.039083,0 .050163,0.010598,-0.006345,-0.000692,0.008181,0.005103,-0.001624,0.016935,-0.019326,-0.007844,0.029451,-0.001724,-0.008996,0.003674,0.005028,-0.007128,0.010606,0.005193,-0.01861,-0.006507,0.008023
FRA_IA:BES1248,0.12862,0.127957,0.064488,0.042636, 0.043085,0.027052,0.007755,-0.003461,0.001636,0.007836,0.010718,0.011839,-0.00892,-0.007569,0.016015,-0.000663,-0.001434,0.004687,-0.015461,0.004002,0.01984,-0.002597,-0.009367,0.008555,-0.016765
average,0.1308965,0.1335425,0.057511,0.0408595,0.0 46624,0.018825,0.000705,-0.0020765,0.0049085,0.0064695,0.004547,0.014387,-0.014123,-0.0077065,0.022733,-0.0011935,-0.005215,0.0041805,-0.0052165,-0.001563,0.015223,0.001298,-0.0139885,0.001024,-0.004371

Distance to: average
0.02910918 French_Brittany
0.02991663 English
0.03013606 Welsh
0.03041254 Orcadian

^^^ These four (or at least 3 from them) could be Southern Gauls as well, and one of them is also very northern:


Iberia_Northeast_Empuries1:I8203,0.126344,0.14319, 0.051288,0.021641,0.037238,-0.005299,0.00094,0.004846,0.024747,0.026971,-0.003897,0.008992,-0.025272,-0.026011,0.008822,0.004906,0.012126,-0.003547,0.003268,0.003627,0.007487,0.000618,-0.001972,0.004458,-0.00467
Iberia_Northeast_Empuries1:I8206,0.125205,0.127957 ,0.053928,0.048773,0.040931,0.013387,0.006815,0.00 6231,0.012067,0.014214,-0.001786,-0.005245,-0.015163,-0.018029,0.021444,0.011403,0.011735,0.010262,0.003 897,0.009129,0.009733,-0.000989,-0.004807,0.016629,0.006227
Iberia_Northeast_Empuries1:I8209,0.12862,0.145221, 0.054682,0.013243,0.043393,0.009761,-0.001175,0.006,0.026384,0.033167,-0.006658,0.018434,-0.016501,-0.023121,0.011401,0.004508,0.005085,0.000127,0.003 897,-0.009254,0.001123,0.00507,0.002218,-0.009881,0.008263
Iberia_Northeast_Empuries1:I8214,0.130897,0.138112 ,0.058454,0.026809,0.046162,0.008367,-0.001175,-0.004154,0.016771,0.030069,-0.001624,0.009741,-0.022299,-0.014588,0.010586,0.00769,0.002868,0.002787,0.0036 45,0.001376,0.011105,0.006801,-0.00986,-0.012893,0.005628

On the other hand, these two were Greeks of course:

Distance to: Iberia_Northeast_Empuries2:I8215
0.04754349 Italian_Campania
0.04775060 Italian_Apulia
0.04848198 Italian_Basilicata
0.04888284 Italian_Calabria
(...)

Distance to: Iberia_Northeast_Empuries2:I8208
0.05613572 Sicilian_East
0.05715079 Italian_Apulia
0.05753055 Italian_Campania
0.05772763 Italian_Calabria
(...)

[Peterski]

There should be more than just two FRA_IA_south samples:

All these 5 Iron Age samples are from Languedoc (SF_IA2):

https://i.imgur.com/jRvYAPV.jpg

This one is interesting:


Distance to: FRA_IA:PEY53
0.02640458 Orcadian
0.02762308 Welsh
0.02767553 French_Brittany
0.02777884 English
0.02814059 French_Occitanie
0.02853848 Belgian
0.02868151 English_Cornwall
0.02868161 French_Seine-Maritime
0.02884452 Scottish
0.02902589 French_Nord
0.02907210 Spanish_Pais_Vasco
0.02927580 Irish
0.02935877 Swiss_German
0.02975863 French_Auvergne
0.03009721 French_Alsace
0.03027516 Afrikaner
0.03054290 Spanish_Barcelones
0.03056663 French_Pas-de-Calais
0.03074413 French_Paris
0.03087750 Icelandic
0.03095438 Basque_French
0.03123319 Spanish_Navarra
0.03201201 Spanish_Penedes
0.03205288 Dutch
0.03208144 Spanish_Alacant


(...)

I was told by a French guy, quote:

"Migrations inside/within ancient Gaul were common."

So the northern-shifted ones in the South could be migrants from the north, and the southern-shifted ones in the North vice versa (migrants from the south). As well as some of them could have mixed parents (one north Gaulish parent, one south Gaulish parent).

I think if you add samples from Empuries1 cluster (3 of whom were most likely Gauls rather than Celtiberians), you obtain a more realistic South Gaulish average.

[dududud]

The biggest irony is that the native language of France (French) is not from the Gauls and neither from the Franks but from Romance/Italic. :crazy:

Latinisation like in Sardinia (they spoke a basque related language before). But not genetically in the case of France

[XenophobicPrussian]

How does our Prussian Xeno Warrior Prince feel about his beloved Nordic Viking I1-M253 haplogrup being possibly descended from the ancient Frenchies?: :laugh:

https://www.theapricity.com/forum/showthread.php?323831-I1-found-in-Neolithic-France&p=6719591&viewfull=1#post6719591

There's also an I1 in neolithic Croatia/Sweden that we've known about for years. Obviously, all I2/I1 are ultimately from HGs, and all R from eastern/Siberian HGs, don't see why it being present in France would be a source of butthurt. The only question is if the modern Scandinavian clade came with Bell Beakers or is from Scandinavian neolithics, also whether or not the French IA one is the Germanic clade or not.

So yeah, no butthurt, only butthurt I see is from the Frenchies trying to cope with now a 3th confirmed major outside group contributing to the genetics of modern French(Romans, Germanics, and now even a very different population from pre-Celtic French, from the core Hallstatt Alpine area, and no your outright weird genetic analysis methods of using K36 the way you do doesn't change this).

and yes you're right, those Iberians are probably the best proxy for average southern Gauls, not the two northern shifted ones from the paper, south French were probably already similar or identical to Hallstatt Celts even before Celts spread the language there and potentially didn't speak an IE language like parts of Iberia despite having heavy Bell Beaker admixture(although probably a bit more southern shifted because they probably have admixture from the northern outliers, and northern/western Gauls or pre-Celtic French).

[Aren]

I was told by a French guy, quote:

"Migrations inside/within ancient Gaul were common."

So the northern-shifted ones in the South could be migrants from the north, and the southern-shifted ones in the North vice versa (migrants from the south). As well as some of them could have mixed parents (one north Gaulish parent, one south Gaulish parent).

I think if you add samples from Empuries1 cluster (3 of whom were most likely Gauls rather than Celtiberians), you obtain a more realistic South Gaulish average.
I'm hoping the higher coverage samples from this study will clear things up cause right now it looks completely random how the most southern shifted samples are from Alsace and meanwhile three of the most Steppe scoring IA Gauls(FRA_IA:BES1248, FRA_IA: PEY53, Empuries1:I8206) all are from the extreme south.

[Peterski]

Actually I wouldn't be surprised if an Armorican-like or Breton-like population extended further south in Pre-Roman Iron Age, than today.

I think it is a pity that this study did not sample Armorica and many other regions.

Also no any ancient samples from Aquitania.

=====

I used to think that modern Bretons are mainly descended from Britons whose migration displaced Roman era Armoricans, but maybe I was wrong.

In 23andMe for example, Bretons do score "British&Irish" rather than "French&German". But maybe ancient Armoricans would score the same.

[Lucas]

There should me more than just two FRA_IA_south samples:

All these 5 Iron Age samples are from Languedoc (SF_IA2):

https://i.imgur.com/jRvYAPV.jpg

This one is interesting:


Distance to: FRA_IA:PEY53
0.02640458 Orcadian
0.02762308 Welsh
0.02767553 French_Brittany
0.02777884 English
0.02814059 French_Occitanie
0.02853848 Belgian
0.02868151 English_Cornwall
0.02868161 French_Seine-Maritime
0.02884452 Scottish
0.02902589 French_Nord
0.02907210 Spanish_Pais_Vasco
0.02927580 Irish
0.02935877 Swiss_German
0.02975863 French_Auvergne
0.03009721 French_Alsace
0.03027516 Afrikaner
0.03054290 Spanish_Barcelones
0.03056663 French_Pas-de-Calais
0.03074413 French_Paris
0.03087750 Icelandic
0.03095438 Basque_French
0.03123319 Spanish_Navarra
0.03201201 Spanish_Penedes
0.03205288 Dutch
0.03208144 Spanish_Alacant



I was told by a French guy, quote:

"Migrations inside/within ancient Gaul were common."

So the northern-shifted ones in the South could be migrants from the north, and the southern-shifted ones in the North vice versa (migrants from the south). As well as some of them could have mixed parents (one north Gaulish parent, one south Gaulish parent).

I think if you add samples from Empuries1 cluster (3 of whom were most likely Gauls rather than Celtiberians), you obtain a more realistic South Gaulish average.

Davidski in g25 add only two for south. So check in k15 or k36 rest of the southerners. You have them already.

[Peterski]

Davidski sent me this G25-based PCA shortly after he had added them to G25:

(what is visible here is that all of French IA samples fall within modern European variation, while some French BA fall rather outside of modern variation)

https://i.imgur.com/kDz0IDa.png

Here are BA samples which are outside (or almost outside) modern variation:

https://i.imgur.com/P6fJxTL.png

[Lucas]

OK, there is samples spreadshee which was missing at first on the website.
Data on Y-DNA subclades, locations etc.

https://drive.google.com/file/d/1P3iJjYWZd14OFgqlRLDN91VpoFnteFlR/view

[J. Ketch]

Davidski sent me this G25-based PCA shortly after he had added them to G25:

(what is visible here is that all of French IA samples fall within modern European variation, while some French BA fall rather outside of modern variation)

https://i.imgur.com/kDz0IDa.png

Here are BA samples which are outside (or almost outside) modern variation:

https://i.imgur.com/P6fJxTL.png
The Purple IA samples are heading off in a different direction South, towards more pure Farmers, i.e. LBK descended Alpines.

[Lucas]

Davidski in g25 add only two for south. So check in k15 or k36 rest of the southerners. You have them already.

Ok here is only one additional from the south https://www.theapricity.com/forum/showthread.php?321209-Ancient-French-calculator-results

At least Iberian-like in K15.

Distance to: PECH5
16.65402654 Spanish_Galicia
17.08797238 France-South
17.37618485 Spanish_Cantabria
18.19327898 Portuguese
18.65733100 Spanish_Cataluna
18.94331280 Spanish_Castilla_Y_Leon
19.02468659 Southwest_French
19.36992514 Italy_Aostavalley
19.84696198 French
20.07233669 Spanish_Extremadura
20.76111750 Spanish_Murcia
20.90741734 Spanish_Aragon
21.12984146 Spanish_Castilla_La_Mancha
21.42933270 Italy_Piedmont
21.73571255 Spanish_Valencia
22.23409544 Spanish_Andalucia
22.23494322 Italy_Trentino
23.35971104 Italy_Lombardy
23.69396548 South_Dutch
23.75248619 Italy_Veneto
23.81081057 Austria-Tyrol
23.94552150 Southwest_English
24.01722715 German_Bavarian
24.18780891 Belgium
24.46335423 West_German


Target: PECH5
Distance: 16.4448% / 16.44477049 | ADC: 1x
81.8 Spanish_Galicia
18.2 Spanish_Cantabria

Target: PECH5
Distance: 15.7704% / 15.77042478 | ADC: 0.5x
56.4 Spanish_Galicia
27.0 Spanish_Cantabria
10.6 English_Midlands
6.0 Sardinian

Target: PECH5
Distance: 13.0902% / 13.09019922 | ADC: 0.25x
59.6 English_Midlands
40.0 Sardinian
0.4 French_Basque

Target: PECH5
Distance: 13.0183% / 13.01826143
41.6 Sardinian
40.2 West_Norwegian
10.2 English_Midlands
7.8 French_Basque
0.2 Karitiana

[gixajo]

Iberia_Northeast_Empuries1:I8203,0.126344,0.14319, 0.051288,0.021641,0.037238,-0.005299,0.00094,0.004846,0.024747,0.026971,-0.003897,0.008992,-0.025272,-0.026011,0.008822,0.004906,0.012126,-0.003547,0.003268,0.003627,0.007487,0.000618,-0.001972,0.004458,-0.00467
Iberia_Northeast_Empuries1:I8206,0.125205,0.127957 ,0.053928,0.048773,0.040931,0.013387,0.006815,0.00 6231,0.012067,0.014214,-0.001786,-0.005245,-0.015163,-0.018029,0.021444,0.011403,0.011735,0.010262,0.003 897,0.009129,0.009733,-0.000989,-0.004807,0.016629,0.006227
Iberia_Northeast_Empuries1:I8209,0.12862,0.145221, 0.054682,0.013243,0.043393,0.009761,-0.001175,0.006,0.026384,0.033167,-0.006658,0.018434,-0.016501,-0.023121,0.011401,0.004508,0.005085,0.000127,0.003 897,-0.009254,0.001123,0.00507,0.002218,-0.009881,0.008263
Iberia_Northeast_Empuries1:I8214,0.130897,0.138112 ,0.058454,0.026809,0.046162,0.008367,-0.001175,-0.004154,0.016771,0.030069,-0.001624,0.009741,-0.022299,-0.014588,0.010586,0.00769,0.002868,0.002787,0.0036 45,0.001376,0.011105,0.006801,-0.00986,-0.012893,0.005628
FRA_IA:ATT26,0.1161,0.132019,0.044877,0.039083,0.0 38776,0.013108,-0.006815,0.011999,0.014112,0.020957,-0.001299,0.007793,-0.024678,-0.011836,0.007872,0.007027,0.006128,0.008361,0.010 307,-0.013632,0.003993,0.00779,0.020706,0.000482,0.0087 42
FRA_IA:BFM265,0.122929,0.148267,0.05242,0.037468,0 .047086,0.018128,-0.00235,-0.000231,0.008795,0.012392,-0.007957,0.005695,-0.015312,-0.007844,0.020765,-0.003182,-0.016428,0.009122,0.006034,0.007504,-0.002246,0.000124,0.001109,-0.00012,0.002036
FRA_IA:COL11,0.122929,0.13405,0.058454,0.064923,0. 037238,0.020917,-0.008225,0.01223,0.008795,0.011116,-0.002273,0.002548,-0.016353,-0.003991,0.017508,-0.008221,-0.014603,-0.005954,0.006662,0.005378,0.006489,0.008161,0.010 106,-0.001446,0.000599
FRA_IA:COL153A,0.130897,0.158423,0.045254,0.02584, 0.036622,0.012271,0.006815,0.008538,0.003272,0.015 126,0.009419,-0.007194,0.001189,-0.000963,0.005022,0.003182,-0.013038,0.00038,0.016592,0.001,-0.015847,0.007543,0.000863,-0.018798,0.005389
FRA_IA:COL153i,0.120652,0.127957,0.046763,0.024548 ,0.045239,-0.006136,0.015276,0.004154,0.015748,0.034078,-0.00341,0.017235,-0.022448,-0.010459,0.012622,-0.026518,0.005476,0.008868,-0.001131,0.015883,-0.000749,-0.000742,-0.010476,0.003253,0.008143
FRA_IA:ERS86,0.122929,0.161469,0.040729,0.030039,0 .040623,0.008646,-0.010105,0.005077,0.006954,0.007289,-0.00065,0.005845,-0.010852,-0.002752,0.005157,0.021745,-0.004172,0.009755,0.011313,-0.005753,0.000624,0.012365,-0.004683,-0.012893,0.00012
FRA_IA:ERS88,0.130897,0.155376,0.044123,0.005168,0 .033237,0.000837,-0.004935,-0.005769,0.011249,0.02442,-0.004872,-0.001499,-0.023042,-0.016239,0.012079,-0.017104,-0.014603,-0.006461,-0.014204,-0.002251,-0.002496,0.000742,0.001356,0,0.000239
FRA_IA:ERS1164,0.124067,0.121864,0.052797,0.030039 ,0.033237,0.010877,0.011516,0.00923,0,0.019135,-0.002598,-0.01154,-0.01115,-0.012799,0.009908,0.006364,-0.002738,0.00038,0.001257,-0.004627,-0.007487,0.003957,0.002218,-0.011809,0.005389
FRA_IA:Jeb8,0.118376,0.138112,0.05506,0.027132,0.0 45239,-0.008367,-0.00329,0.003461,0.012271,0.010023,-0.00341,0.007194,-0.020812,-0.010046,0.016694,0.007425,-0.000391,-0.001647,0.000377,0.005378,-0.003868,-0.00136,-0.009983,-0.003615,0.007065
FRA_IA:NOR2B6,0.120652,0.147252,0.044877,0.005814, 0.0397,0.003347,-0.010105,0.007384,0.016975,0.014943,0.010718,0.011 09,-0.008771,-0.004542,0.012215,0.018165,0.027772,0.004941,0.005 405,0.002876,0.006364,-0.007543,-0.001109,-0.00253,-0.007784
FRA_IA:NOR3-6,0.133173,0.146236,0.050157,0.046189,0.029852,0.0 17291,-0.007755,0.005077,0.012885,0.001276,-0.002598,0.015137,-0.022745,-0.013212,0.020087,-0.007823,-0.021383,0.007095,-0.001006,0.006753,0.004866,0.005193,-0.002835,-0.007953,-0.000239
FRA_IA:NOR3-15,0.117238,0.149283,0.061471,0.022287,0.043393,-0.005299,0.005405,-0.001615,0.013703,0.018224,-0.005196,0.00045,-0.014569,-0.012111,0.022394,-0.005171,-0.019558,-0.006841,-0.004902,0.013381,-0.002121,-0.002844,-0.015652,-0.003615,0.002395
FRA_IA:NOR4,0.12862,0.151314,0.058454,0.001615,0.0 46778,-0.006136,-0.000705,0.014538,0.023929,0.031345,-0.013153,0.006594,-0.022448,-0.016377,-0.016965,0.004773,0.005737,-0.010135,0.011941,0.005503,0.006239,0.006183,-0.011462,0.007471,-0.002994
FRA_IA:PEY53,0.133173,0.139128,0.050534,0.039083,0 .050163,0.010598,-0.006345,-0.000692,0.008181,0.005103,-0.001624,0.016935,-0.019326,-0.007844,0.029451,-0.001724,-0.008996,0.003674,0.005028,-0.007128,0.010606,0.005193,-0.01861,-0.006507,0.008023
FRA_IA:BES1248,0.12862,0.127957,0.064488,0.042636, 0.043085,0.027052,0.007755,-0.003461,0.001636,0.007836,0.010718,0.011839,-0.00892,-0.007569,0.016015,-0.000663,-0.001434,0.004687,-0.015461,0.004002,0.01984,-0.002597,-0.009367,0.008555,-0.016765

Target: gixajo_scaled
Distance: 2.4232% / 0.02423216
25.0 FRA_IA:ERS88
22.0 FRA_IA:NOR2B6
19.6 Iberia_Northeast_Empuries1:I8209
16.6 FRA_IA:NOR3-15
12.2 FRA_IA:COL153i
4.6 FRA_IA:NOR4

Distance to: gixajo_scaled
0.03928515 Iberia_Northeast_Empuries1:I8209
0.04189850 Iberia_Northeast_Empuries1:I8203
0.04212763 FRA_IA:ERS88
0.04416381 FRA_IA:NOR3-15
0.04483023 FRA_IA:Jeb8
0.04494347 Iberia_Northeast_Empuries1:I8214
0.05128186 FRA_IA:NOR2B6
0.05148714 FRA_IA:COL153i
0.05272917 FRA_IA:NOR4
0.05512703 FRA_IA:BFM265
0.05940530 FRA_IA:ERS86
0.06212623 FRA_IA:ERS1164
0.06278213 FRA_IA:COL153A
0.06312113 FRA_IA:PEY53
0.06567193 FRA_IA:ATT26
0.06938475 FRA_IA:NOR3-6
0.07213493 Iberia_Northeast_Empuries1:I8206
0.07652085 FRA_IA:BES1248
0.08207717 FRA_IA:COL11

[dududud]

The provenance of these samples:
https://drive.google.com/file/d/1P3iJjYWZd14OFgqlRLDN91VpoFnteFlR/view

[Defcon2]

Iberia_Northeast_Empuries1:I8203,0.126344,0.14319, 0.051288,0.021641,0.037238,-0.005299,0.00094,0.004846,0.024747,0.026971,-0.003897,0.008992,-0.025272,-0.026011,0.008822,0.004906,0.012126,-0.003547,0.003268,0.003627,0.007487,0.000618,-0.001972,0.004458,-0.00467
Iberia_Northeast_Empuries1:I8206,0.125205,0.127957 ,0.053928,0.048773,0.040931,0.013387,0.006815,0.00 6231,0.012067,0.014214,-0.001786,-0.005245,-0.015163,-0.018029,0.021444,0.011403,0.011735,0.010262,0.003 897,0.009129,0.009733,-0.000989,-0.004807,0.016629,0.006227
Iberia_Northeast_Empuries1:I8209,0.12862,0.145221, 0.054682,0.013243,0.043393,0.009761,-0.001175,0.006,0.026384,0.033167,-0.006658,0.018434,-0.016501,-0.023121,0.011401,0.004508,0.005085,0.000127,0.003 897,-0.009254,0.001123,0.00507,0.002218,-0.009881,0.008263
Iberia_Northeast_Empuries1:I8214,0.130897,0.138112 ,0.058454,0.026809,0.046162,0.008367,-0.001175,-0.004154,0.016771,0.030069,-0.001624,0.009741,-0.022299,-0.014588,0.010586,0.00769,0.002868,0.002787,0.0036 45,0.001376,0.011105,0.006801,-0.00986,-0.012893,0.005628
FRA_IA:ATT26,0.1161,0.132019,0.044877,0.039083,0.0 38776,0.013108,-0.006815,0.011999,0.014112,0.020957,-0.001299,0.007793,-0.024678,-0.011836,0.007872,0.007027,0.006128,0.008361,0.010 307,-0.013632,0.003993,0.00779,0.020706,0.000482,0.0087 42
FRA_IA:BFM265,0.122929,0.148267,0.05242,0.037468,0 .047086,0.018128,-0.00235,-0.000231,0.008795,0.012392,-0.007957,0.005695,-0.015312,-0.007844,0.020765,-0.003182,-0.016428,0.009122,0.006034,0.007504,-0.002246,0.000124,0.001109,-0.00012,0.002036
FRA_IA:COL11,0.122929,0.13405,0.058454,0.064923,0. 037238,0.020917,-0.008225,0.01223,0.008795,0.011116,-0.002273,0.002548,-0.016353,-0.003991,0.017508,-0.008221,-0.014603,-0.005954,0.006662,0.005378,0.006489,0.008161,0.010 106,-0.001446,0.000599
FRA_IA:COL153A,0.130897,0.158423,0.045254,0.02584, 0.036622,0.012271,0.006815,0.008538,0.003272,0.015 126,0.009419,-0.007194,0.001189,-0.000963,0.005022,0.003182,-0.013038,0.00038,0.016592,0.001,-0.015847,0.007543,0.000863,-0.018798,0.005389
FRA_IA:COL153i,0.120652,0.127957,0.046763,0.024548 ,0.045239,-0.006136,0.015276,0.004154,0.015748,0.034078,-0.00341,0.017235,-0.022448,-0.010459,0.012622,-0.026518,0.005476,0.008868,-0.001131,0.015883,-0.000749,-0.000742,-0.010476,0.003253,0.008143
FRA_IA:ERS86,0.122929,0.161469,0.040729,0.030039,0 .040623,0.008646,-0.010105,0.005077,0.006954,0.007289,-0.00065,0.005845,-0.010852,-0.002752,0.005157,0.021745,-0.004172,0.009755,0.011313,-0.005753,0.000624,0.012365,-0.004683,-0.012893,0.00012
FRA_IA:ERS88,0.130897,0.155376,0.044123,0.005168,0 .033237,0.000837,-0.004935,-0.005769,0.011249,0.02442,-0.004872,-0.001499,-0.023042,-0.016239,0.012079,-0.017104,-0.014603,-0.006461,-0.014204,-0.002251,-0.002496,0.000742,0.001356,0,0.000239
FRA_IA:ERS1164,0.124067,0.121864,0.052797,0.030039 ,0.033237,0.010877,0.011516,0.00923,0,0.019135,-0.002598,-0.01154,-0.01115,-0.012799,0.009908,0.006364,-0.002738,0.00038,0.001257,-0.004627,-0.007487,0.003957,0.002218,-0.011809,0.005389
FRA_IA:Jeb8,0.118376,0.138112,0.05506,0.027132,0.0 45239,-0.008367,-0.00329,0.003461,0.012271,0.010023,-0.00341,0.007194,-0.020812,-0.010046,0.016694,0.007425,-0.000391,-0.001647,0.000377,0.005378,-0.003868,-0.00136,-0.009983,-0.003615,0.007065
FRA_IA:NOR2B6,0.120652,0.147252,0.044877,0.005814, 0.0397,0.003347,-0.010105,0.007384,0.016975,0.014943,0.010718,0.011 09,-0.008771,-0.004542,0.012215,0.018165,0.027772,0.004941,0.005 405,0.002876,0.006364,-0.007543,-0.001109,-0.00253,-0.007784
FRA_IA:NOR3-6,0.133173,0.146236,0.050157,0.046189,0.029852,0.0 17291,-0.007755,0.005077,0.012885,0.001276,-0.002598,0.015137,-0.022745,-0.013212,0.020087,-0.007823,-0.021383,0.007095,-0.001006,0.006753,0.004866,0.005193,-0.002835,-0.007953,-0.000239
FRA_IA:NOR3-15,0.117238,0.149283,0.061471,0.022287,0.043393,-0.005299,0.005405,-0.001615,0.013703,0.018224,-0.005196,0.00045,-0.014569,-0.012111,0.022394,-0.005171,-0.019558,-0.006841,-0.004902,0.013381,-0.002121,-0.002844,-0.015652,-0.003615,0.002395
FRA_IA:NOR4,0.12862,0.151314,0.058454,0.001615,0.0 46778,-0.006136,-0.000705,0.014538,0.023929,0.031345,-0.013153,0.006594,-0.022448,-0.016377,-0.016965,0.004773,0.005737,-0.010135,0.011941,0.005503,0.006239,0.006183,-0.011462,0.007471,-0.002994
FRA_IA:PEY53,0.133173,0.139128,0.050534,0.039083,0 .050163,0.010598,-0.006345,-0.000692,0.008181,0.005103,-0.001624,0.016935,-0.019326,-0.007844,0.029451,-0.001724,-0.008996,0.003674,0.005028,-0.007128,0.010606,0.005193,-0.01861,-0.006507,0.008023
FRA_IA:BES1248,0.12862,0.127957,0.064488,0.042636, 0.043085,0.027052,0.007755,-0.003461,0.001636,0.007836,0.010718,0.011839,-0.00892,-0.007569,0.016015,-0.000663,-0.001434,0.004687,-0.015461,0.004002,0.01984,-0.002597,-0.009367,0.008555,-0.016765

Target: gixajo_scaled
Distance: 2.4232% / 0.02423216
25.0 FRA_IA:ERS88
22.0 FRA_IA:NOR2B6
19.6 Iberia_Northeast_Empuries1:I8209
16.6 FRA_IA:NOR3-15
12.2 FRA_IA:COL153i
4.6 FRA_IA:NOR4

Distance to: gixajo_scaled
0.03928515 Iberia_Northeast_Empuries1:I8209
0.04189850 Iberia_Northeast_Empuries1:I8203
0.04212763 FRA_IA:ERS88
0.04416381 FRA_IA:NOR3-15
0.04483023 FRA_IA:Jeb8
0.04494347 Iberia_Northeast_Empuries1:I8214
0.05128186 FRA_IA:NOR2B6
0.05148714 FRA_IA:COL153i
0.05272917 FRA_IA:NOR4
0.05512703 FRA_IA:BFM265
0.05940530 FRA_IA:ERS86
0.06212623 FRA_IA:ERS1164
0.06278213 FRA_IA:COL153A
0.06312113 FRA_IA:PEY53
0.06567193 FRA_IA:ATT26
0.06938475 FRA_IA:NOR3-6
0.07213493 Iberia_Northeast_Empuries1:I8206
0.07652085 FRA_IA:BES1248
0.08207717 FRA_IA:COL11

Target: Juan_scaled
Distance: 4.5026% / 0.04502593
42.8 FRA_IA:ERS88
24.4 FRA_IA:COL153i
21.6 FRA_IA:NOR4
11.2 FRA_IA:ATT26


Distance to: Juan_scaled
0.05372398 FRA_IA:ERS88
0.05865324 Iberia_Northeast_Empuries1:I8209
0.05978581 Iberia_Northeast_Empuries1:I8203
0.06079094 FRA_IA:COL153i
0.06325204 FRA_IA:NOR4
0.06580361 FRA_IA:NOR3-15
0.06682850 FRA_IA:Jeb8
0.06819813 Iberia_Northeast_Empuries1:I8214
0.07080622 FRA_IA:ATT26
0.07187085 FRA_IA:BFM265
0.07346020 FRA_IA:NOR2B6
0.07527434 FRA_IA:ERS1164
0.08049998 FRA_IA:ERS86
0.08278253 FRA_IA:COL153A
0.08411912 FRA_IA:PEY53
0.08563772 FRA_IA:NOR3-6
0.08771873 Iberia_Northeast_Empuries1:I8206
0.09166822 FRA_IA:BES1248
0.09278370 FRA_IA:COL11

[gixajo]

Target: Juan_scaled
Distance: 4.5026% / 0.04502593
42.8 FRA_IA:ERS88
24.4 FRA_IA:COL153i
21.6 FRA_IA:NOR4
11.2 FRA_IA:ATT26


Distance to: Juan_scaled
0.05372398 FRA_IA:ERS88
0.05865324 Iberia_Northeast_Empuries1:I8209
0.05978581 Iberia_Northeast_Empuries1:I8203
0.06079094 FRA_IA:COL153i
0.06325204 FRA_IA:NOR4
0.06580361 FRA_IA:NOR3-15
0.06682850 FRA_IA:Jeb8
0.06819813 Iberia_Northeast_Empuries1:I8214
0.07080622 FRA_IA:ATT26
0.07187085 FRA_IA:BFM265
0.07346020 FRA_IA:NOR2B6
0.07527434 FRA_IA:ERS1164
0.08049998 FRA_IA:ERS86
0.08278253 FRA_IA:COL153A
0.08411912 FRA_IA:PEY53
0.08563772 FRA_IA:NOR3-6
0.08771873 Iberia_Northeast_Empuries1:I8206
0.09166822 FRA_IA:BES1248
0.09278370 FRA_IA:COL11


No me cuadra nada como están clasificados esos samples. Ni los supuestamente franceses ni los supuestamente ibéricos.

[gixajo]

Repeated.

[SharpFork]

I don't want to start the same discussion from 0, but I still don't exactly see yet the smoking gun evidence for the Hallstatt = Southern French theory(but I mean the discussion from my point of view is incredibly specific, Central-North French-like vs South French-like), there is still an incredible amount of variation:

Of the samples Lukasz posted:

1 of the 2 northern ones is British-like, the other is North French like(Paris)

The central ones show a lot of difference:

British(Cornish)
2 South German(Alsace)
4 Southern French(all Occitanie)
2 North-East Iberian(roughly Catalan regions)
2 Northern(Breton and Belgian)

Outside generalized statements, this is a brief view of what I see when separating by locale and time:

Northern Alsace(Nordhouse): The 2 early samples from the 8th century BCE are South-Central French like(Occitanie and Swiss French), the 2 later one from around the 6th century BCE are very different, one is Breton like and the other is North-East Spanish like.

Northern Alsace(Erstein): Those 3 samples from 400-100 BCE are also very diverse, one is South German like, one is Belgian like and another is NE Spanish like.

Southern Alsace(Sainte-Croix-en-Plaine): The single sample from 600-400 BCE is South French like(Occitanie like always)

Southern Alsace(Colmar): The 3 samples from the late 4th century BCE show diversity like always: one is Orcadian-like, one is South German and another South French.

Picardy(Attichy & Bucy-le-Long): one sample is from around 350-200 BCE and the other from the late 3th century BCE, the former is North French-like(Paris) and the later is British like(Cornwall)

I can't really make sense of this myself, I need to look at the Bronze age samples too.

[XenophobicPrussian]

I don't want to start the same discussion from 0, but I still don't exactly see yet the smoking gun evidence for the Hallstatt = Southern French theory(but I mean the discussion from my point of view is incredibly specific, Central-North French-like vs South French-like), there is still an incredible amount of variation:

Of the samples Lukasz posted:

1 of the 2 northern ones is British-like, the other is North French like(Paris)

The central ones show a lot of difference:

British(Cornish)
2 South German(Alsace)
4 Southern French(all Occitanie)
2 North-East Iberian(roughly Catalan regions)
2 Northern(Breton and Belgian)

Outside generalized statements, this is a brief view of what I see when separating by locale and time:

Northern Alsace(Nordhouse): The 2 early samples from the 8th century BCE are South-Central French like(Occitanie and Swiss French), the 2 later one from around the 6th century BCE are very different, one is Breton like and the other is North-East Spanish like.

Northern Alsace(Erstein): Those 3 samples from 400-100 BCE are also very diverse, one is South German like, one is Belgian like and another is NE Spanish like.

Southern Alsace(Sainte-Croix-en-Plaine): The single sample from 600-400 BCE is South French like(Occitanie like always)

Southern Alsace(Colmar): The 3 samples from the late 4th century BCE show diversity like always: one is Orcadian-like, one is South German and another South French.

Picardy(Attichy & Bucy-le-Long): one sample is from around 350-200 BCE and the other from the late 3th century BCE, the former is North French-like(Paris) and the later is British like(Cornwall)

I can't really make sense of this myself, I need to look at the Bronze age samples too.
I don't see how you get to any conclusion other than Hallstatt Celts were southern French/Iberian like.

Yes, there's an incredible amount of variation, that's the entire reason why Hallstatt Celts are going to be southern French/Iberian like. Levels of that kind of variation in steppe/neolithic farmer ancestry will not survive from the early Bronze Age to the Iron Age, that's just not going to happen, not without some sort of rigid caste system, and I'm not even sure the differences between Indian castes are that large. This means the variation is explained by locals, mixed people, and immigrants. The Breton/British clustering ones are clearly the locals, the others mixes(could be just more southernly located locals too though), the southern shifted ones are quite clearly immigrants from the Hallstatt zone(potentially from southern France, but there's literally no reason to think this as we already have the southern shifted population appearing in the Alpine Hallstatt Zone, and Switzerland/Austria are closer to Alsace than southern France, and there's actually known migration and cultural diffusion from the Alpine area to all surrounding areas, unlike from southern France).

Right now the only IA sample from Switzerland is Iberian clustering(for the record, N. Iberian to S. France clustering can just be natural variation in a population, but N. Iberian to Belgian? not happening), and you don't even need more as this paper pretty much ends the argument. More IA samples from Switzerland/Austria would be great but as far as I'm concerned this is settled.

[Peterski]

That is not an "incredible" amount of variation, that is exactly the same amount of variation as among modern French population. And academic samples show that, also today, this variation does not strictly correlate with geography. There are people belonging to different genetic clusters but living in the same area - this is also the case in modern France based on academic samples.

Apparently it was the case in ancient Gaul too.

And yes, studies on the modern French have been criticized for not paying enough attention to origins of grandparents of the sampled individuals. But it is exactly the same case with ancient samples - we know nothing about genealogy of grandparents of ATT26 (for example). Did they at least examine the strontium isotopes to see who was locally born?

[SharpFork]

That is not an "incredible" amount of variation, that is exactly the same amount of variation as among modern French population. And academic samples show that, also today, this variation does not strictly correlate with geography. There are people belonging to different genetic clusters but living in the same area - this is also the case in modern France based on academic samples.

Apparently it was the case in ancient Gaul too.

And yes, studies on the modern French have been criticized for not paying enough attention to origins of grandparents of the sampled individuals. But it is exactly the same case with ancient samples - we know nothing about genealogy of grandparents of ATT26 (for example). Did they at least examine the strontium isotopes to see who was locally born?
Do we have the new Bronze age samples in G25?

[Lucas]

Do we have the new Bronze age samples in G25?

Not in official dataset. Davidski published them on forum Anthrogenica only as there have shit coverage (the same for IA).



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[SharpFork]

I don't see how you get to any conclusion other than Hallstatt Celts were southern French/Iberian like.

Yes, there's an incredible amount of variation, that's the entire reason why Hallstatt Celts are going to be southern French/Iberian like. Levels of that kind of variation in steppe/neolithic farmer ancestry will not survive from the early Bronze Age to the Iron Age, that's just not going to happen, not without some sort of rigid caste system, and I'm not even sure the differences between Indian castes are that large. This means the variation is explained by locals, mixed people, and immigrants. The Breton/British clustering ones are clearly the locals, the others mixes(could be just more southernly located locals too though), the southern shifted ones are quite clearly immigrants from the Hallstatt zone(potentially from southern France, but there's literally no reason to think this as we already have the southern shifted population appearing in the Alpine Hallstatt Zone, and Switzerland/Austria are closer to Alsace than southern France, and there's actually known migration and cultural diffusion from the Alpine area to all surrounding areas, unlike from southern France).

Right now the only IA sample from Switzerland is Iberian clustering(for the record, N. Iberian to S. France clustering can just be natural variation in a population, but N. Iberian to Belgian? not happening), and you don't even need more as this paper pretty much ends the argument. More IA samples from Switzerland/Austria would be great but as far as I'm concerned this is settled.
I'll examine the Bronze age samples later but at least the Bronze age and Iron Age averages are still very close so the most evident change between the 2 periods is a reduction of variation, not a particular southern shift, but like I said I'll look into it more, Alsace especially.

A bit off-topic, but you previously mentioned that the original proto-Celts would have been quite Northern before reaching the South German region, what do you believe the general chronology of events was? Where and when did the Proto-Celts come from and when did they mix with the Southern-looking Danubian/Upper Rhine populations?