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Macedonian Y-DNA
Y-Chromosome Haplogroup Distribution – Vardar Region
Sample size (N) = 120
1. Major Haplogroup Composition
Haplogroup Count Percentage
E 36 30.00%
I 28 23.33%
J 24 20.00%
R1b 21 17.50%
R1a 10 8.33%
L 1 0.83%
TOTAL 120 100.00%
2. Detailed Haplogroup & Subclade Breakdown
Haplogroup E – 36 (30.00%)
E-V13 Cluster – 29 (24.17%)
Subclade Count Percentage
E-V13 (unspecified) 26 21.67%
E-V13 (regional / unresolved) 3 2.50%
E-V13 subtotal 29 24.17%
Other E Lineages – 7 (5.83%)
Subclade Count Percentage
E-V22 2 1.67%
E-L677 2 1.67%
E-CTS2548 1 0.83%
E-PH1173 1 0.83%
E-M10 (E-V68) 1 0.83%
Other E subtotal 7 5.83%
E TOTAL: 36 (30.00%)
Haplogroup I – 28 (23.33%)
I2 Lineages – 26 (21.67%)
Subclade Count Percentage
I2a (unspecified) 12 10.00%
I-CTS10057 (incl. CTS10100, Z162) 6 5.00%
I-CTS5966 4 3.33%
I2-Y3120 (incl. PH908, S17250) 4 3.33%
I2 subtotal 26 21.67%
Other I – 2 (1.67%)
Subclade Count Percentage
I (unspecified) 2 1.67%
I TOTAL: 28 (23.33%)
Haplogroup J – 24 (20.00%)
J2 Lineages – 21 (17.50%)
Subclade Count Percentage
J2a (unspecified) 13 10.83%
J2a-L24 1 0.83%
J-L70 (J2a) 1 0.83%
J-M67 (J2a) 3 2.50%
J2b-M241 / Z38299 3 2.50%
J2 subtotal 21 17.50%
J1 – 3 (2.50%)
Subclade Count Percentage
J1 3 2.50%
J TOTAL: 24 (20.00%)
Haplogroup R1b – 21 (17.50%)
R1b-Z2013 – 18 (15.00%)
Subclade Count Percentage
R1b-Z2013 18 15.00%
R1b-L421 Cluster – 3 (2.50%)
Subclade Count Percentage
R1b-L421 / downstream 3 2.50%
R1b TOTAL: 21 (17.50%)
Haplogroup R1a – 10 (8.33%)
Subclade Count Percentage
R1a (unspecified) 7 5.83%
R-Z93 (R1a) 1 0.83%
R1a (regional clusters) 2 1.67%
R1a TOTAL: 10 (8.33%)
Haplogroup L – 1 (0.83%)
Subclade Count Percentage
L-L1307 1 0.83%
L TOTAL: 1 (0.83%)
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SKOPJE REGION — Y-DNA Haplogroup Distribution
Total samples (N = 120)
1. Overall Haplogroup Frequencies
Haplogroup Count Percentage
I 33 27.50%
E 30 25.00%
J 24 20.00%
R1b 21 17.50%
R1a 10 8.33%
G 2 1.67%
TOTAL 120 100.00%
2. Haplogroup E — 30 (25.00%)
Subclade Count Percentage
E1b1b1a (xV13) 2 1.67%
E-V13 (unspecified) 23 19.17%
E-V13 → Z1057 3 2.50%
E-V13 → Z5018 1 0.83%
E-V13 → Z5017 1 0.83%
E TOTAL 30 25.00%
3. Haplogroup J — 24 (20.00%)
J2a — 14 (11.67%)
Subclade Count Percentage
J2a (unspecified) 4 3.33%
J-M67 / J2a1b / CTS6372 4 3.33%
J-L24 3 2.50%
J-M410 (xL24) 1 0.83%
J-M92 → PF7412 1 0.83%
J-L26 1 0.83%
J2a TOTAL 14 11.67%
J2b — 9 (7.50%)
Subclade Count Percentage
J2b (unspecified) 4 3.33%
J2b-L283 3 2.50%
J2b-M205 → Y22059 2 1.67%
J2b TOTAL 9 7.50%
J1 — 1 (0.83%)
Subclade Count Percentage
J1-M267 1 0.83%
J TOTAL: 24 (20.00%)
4. Haplogroup I — 33 (27.50%)
I1 — 3 (2.50%)
Subclade Count Percentage
I1 (unspecified) 1 0.83%
I1-P109 → FGC22045 2 1.67%
I1 TOTAL 3 2.50%
I2 — 30 (25.00%)
Subclade Count Percentage
I2a (unspecified) 3 2.50%
I2-CTS5966 1 0.83%
I2-CTS10228 3 2.50%
I2-Y3120 (total) 23 19.17%
I2-Y3120 Breakdown
Subclade Count Percentage
Y3120 (unspecified) 7 5.83%
S17250 (xPH908) 4 3.33%
S17250 → PH908 3 2.50%
Y4884 3 2.50%
Z17855 6 5.00%
Y3120 TOTAL 23 19.17%
I TOTAL: 33 (27.50%)
5. Haplogroup R1a — 10 (8.33%)
Subclade Count Percentage
R1a (unspecified) 6 5.00%
R1a-Z280 → Y2902 2 1.67%
R1a-M458 → L1029 2 1.67%
R1a TOTAL 10 8.33%
6. Haplogroup R1b — 21 (17.50%)
Subclade Count Percentage
R1b-M269 → Z2013 12 10.00%
R1b-L277 3 2.50%
R1b-L421 3 2.50%
R1b-L23 (xZ2103) 2 1.67%
R1b-M269 → L51 → P311 1 0.83%
R1b TOTAL 21 17.50%
7. Haplogroup G — 2 (1.67%)
Subclade Count Percentage
G2a-L497 → L42 1 0.83%
G2a-L497 → L42 → Y128028 1 0.83%
G TOTAL 2 1.67%
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POLOG REGION — Y-DNA Haplogroup Distribution
Total samples (N = 110)
1. Major Haplogroups
Haplogroup Count Percentage
J 34 30.91%
E 23 20.91%
I 23 20.91%
R1a 14 12.73%
R1b 13 11.82%
T 2 1.82%
G 1 0.91%
TOTAL 110 100.00%
2. Haplogroup J — 34 (30.91%)
J2a — 13 (11.82%)
Subclade Count Percentage
J2a-L70 (incl. L397, L398 and derived) 9 8.18%
J2a (unspecified) 4 3.64%
J2a TOTAL 13 11.82%
J2b — 20 (18.18%)
Subclade Count Percentage
J2b (unspecified) 9 8.18%
J2b-L283 (incl. FT134628, Z631) 8 7.27%
J2b-M241 3 2.73%
J2b TOTAL 20 18.18%
J1 — 1 (0.91%)
Subclade Count Percentage
J1-P58 / J1e 1 0.91%
3. Haplogroup E — 23 (20.91%)
Subclade Count Percentage
E-V13 (all downstream clades) 20 18.18%
E-CTS1273 (incl. CTS2374, CTS5371) 2 1.82%
E-L241 1 0.91%
E TOTAL 23 20.91%
4. Haplogroup I — 23 (20.91%)
Subclade Count Percentage
I2-Y3120 (all) 21 19.09%
└─ I2-S17250 → PH908 6 5.45%
I2c-L596 1 0.91%
I2a (unspecified) 1 0.91%
I TOTAL 23 20.91%
5. Haplogroup R1a — 14 (12.73%)
Subclade Count Percentage
R1a-M458 → L1029 2 1.82%
R1a-M417 / M512 8 7.27%
R1a (unspecified) 4 3.64%
R1a TOTAL 14 12.73%
6. Haplogroup R1b — 13 (11.82%)
Subclade Count Percentage
R1b-Z2013 (incl. Y32147) 6 5.45%
R1b-CTS9219 2 1.82%
R1b-P312 1 0.91%
R1b-Z2103 → Z2705 2 1.82%
R1b-PF7558 / PF7562 2 1.82%
R1b TOTAL 13 11.82%
7. Minor Haplogroups
Haplogroup Count Percentage
T-Y11151 2 1.82%
G2a-M406 1 0.91%
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NORTHEAST REGION — Y-DNA Haplogroup Distribution
Total samples (N = 110)
1. Major Haplogroups
Haplogroup Count Percentage
I 34 30.91%
E 28 25.45%
R1b 20 18.18%
J 15 13.64%
R1a 8 7.27%
G 5 4.55%
TOTAL 110 100.00%
2. Haplogroup I — 34 (30.91%)
Subclade Count Percentage
I2-Y3120 (all downstream) 26 23.64%
├─ I2-S17250 → PH908 11 10.00%
I2-CTS11030 1 0.91%
I2-CTS595 1 0.91%
I2-M223 → P78 → Y4884 2 1.82%
I2-S17250 (undifferentiated) 2 1.82%
I2a (unspecified) 2 1.82%
I TOTAL 34 30.91%
3. Haplogroup E — 28 (25.45%)
Subclade Count Percentage
E-V13 (all downstream clades) 22 20.00%
├─ Z1057 / Z5017 / Z5018 (included above) —
E-L241 2 1.82%
E-L250 2 1.82%
E-BY5610 1 0.91%
E-M123 1 0.91%
E TOTAL 28 25.45%
4. Haplogroup R1b — 20 (18.18%)
Subclade Count Percentage
R1b-Z2013 (incl. FT330613) 11 10.00%
R1b-L277 2 1.82%
R1b-Z2103 → Z2705 / Y32147 3 2.73%
R1b-U152 1 0.91%
R1b-R247 2 1.82%
R1b (unspecified M269) 1 0.91%
R1b TOTAL 20 18.18%
5. Haplogroup J — 15 (13.64%)
Subclade Count Percentage
J2b-M205 → Y22059 (incl. Y22063) 11 10.00%
J2a (incl. L70) 2 1.82%
J1-P58 → Z2313 1 0.91%
J2b (unspecified) 1 0.91%
J TOTAL 15 13.64%
6. Haplogroup R1a — 8 (7.27%)
Subclade Count Percentage
R1a-Z280 (incl. YP4278, YP278.2) 4 3.64%
R1a (unspecified) 4 3.64%
R1a TOTAL 8 7.27%
7. Haplogroup G — 5 (4.55%)
Subclade Count Percentage
G2a3 2 1.82%
G2a3b1 1 0.91%
G2a-PF3359 1 0.91%
G-Z2022 1 0.91%
G TOTAL 5 4.55%
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Y-Chromosome Haplogroup Distribution – East Region
Sample size (N = 110)
1. Major Haplogroup Composition
Haplogroup Count Percentage
I 34 30.91%
E 31 28.18%
R1b 22 20.00%
J 16 14.55%
R1a 7 6.36%
TOTAL 110 100.00%
2. Detailed Haplogroup and Subclade Structure
Haplogroup I — 34 (30.91%)
I2 Lineages — 32 (29.09%)
Subclade Count Percentage
I2a (unspecified) 6 5.45%
I2-CTS10228 (incl. CTS5966) 10 9.09%
I2-M423 8 7.27%
I-L68 (I2) 7 6.36%
I2 subtotal 31 28.18%
I1 Lineages — 2 (1.82%)
Subclade Count Percentage
I1-P109 2 1.82%
Haplogroup I TOTAL: 34 (30.91%)
Haplogroup E — 31 (28.18%)
E-V13 Cluster — 29 (26.36%)
Subclade Count Percentage
E-V13 (unspecified) 24 21.82%
E-CTS1273 (CTS2374 / CTS5371) 4 3.64%
E-V13 → CTS5856 → Z5018 → S2979 → FGC11457 1 0.91%
E-V13 subtotal 29 26.36%
Other E Lineages — 2 (1.82%)
Subclade Count Percentage
E (unresolved / non-V13) 2 1.82%
Haplogroup E TOTAL: 31 (28.18%)
Haplogroup R1b — 22 (20.00%)
R1b-Z2103 (Balkan–Carpathian Cluster) — 3 (2.73%)
Subclade Count Percentage
R1b-M269 → L23 → Z2103 → L584 3 2.73%
R1b-Z2013 — 13 (11.81%)
Subclade Count Percentage
R1b-Z2013 13 11.81%
Other R1b Lineages — 6 (5.45%)
Subclade Count Percentage
R1b-Z156 2 1.82%
R-PF7558 (R1b) 1 0.91%
R1b (unspecified) 3 2.73%
Haplogroup R1b TOTAL: 22 (20.00%)
Haplogroup J — 16 (14.55%)
J2 Lineages — 14 (12.73%)
Subclade Count Percentage
J2a-L26 6 5.45%
J-M67 / J2a1b (CTS6372 / PF5120) 3 2.73%
J2a (unspecified) 5 4.55%
Other J Lineages — 2 (1.82%)
Subclade Count Percentage
J2b 1 0.91%
J1-CTS5368 1 0.91%
Haplogroup J TOTAL: 16 (14.55%)
Haplogroup R1a — 7 (6.36%)
Subclade Count Percentage
R1a-L1029 4 3.64%
R1a (unspecified) 3 2.73%
R1a TOTAL 7 6.36%
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Pelagonia Region — Y-Chromosome Haplogroup Distribution
Total Sample Size (N = 120)
1. Major Haplogroup Composition
Haplogroup Count % of Total
E 39 32.50%
I 27 22.50%
J 27 22.50%
R1b 14 11.67%
R1a 9 7.50%
T 3 2.50%
G 1 0.83%
TOTAL 120 100.00%
2. Detailed Haplogroup & Subclade Breakdown
Haplogroup E — 39 (32.50%)
E-V13 Cluster — 33 (27.50%)
Subclade Count % of Total
E-V13 (unspecified) 30 25.00%
E-V13 → CTS5856 → S7461 1 0.83%
E-V13 → CTS5856 → Z5018 → S2979 → Y3183 1 0.83%
E-V13 → CTS5856 → Z5017 → Z5016 → CTS9320 / S26015 1 0.83%
E-V13 subtotal 33 27.50%
Other E Lineages — 6 (5.00%)
Subclade Count % of Total
E-CTS1273 4 3.33%
E-Z1057 2 1.67%
E TOTAL: 39 (32.50%)
Haplogroup I — 27 (22.50%)
I2 Lineages — 23 (19.17%)
Subclade Count % of Total
I2a (unspecified) 9 7.50%
I-M423 6 5.00%
I-S17250 5 4.17%
I-CTS10057 3 2.50%
I2 subtotal 23 19.17%
Other I — 4 (3.33%)
Subclade Count % of Total
I1-L759 2 1.67%
I-Z17855 1 0.83%
I-P78 1 0.83%
I TOTAL: 27 (22.50%)
Haplogroup J — 27 (22.50%)
J2 Lineages — 22 (18.33%)
Subclade Count % of Total
J2a (unspecified) 15 12.50%
J2a-L70 1 0.83%
J2b-L283 4 3.33%
J-M241 (J2b) 1 0.83%
J-FT278595 (J2b) 1 0.83%
J2 subtotal 22 18.33%
J1 — 5 (4.17%)
Subclade Count % of Total
J1 (unspecified) 5 4.17%
J TOTAL: 27 (22.50%)
Haplogroup R1b — 14 (11.67%)
Subclade Count % of Total
R1b-Z2013 11 9.17%
R1b-Z156 1 0.83%
R1b1b2a 1 0.83%
R1b-M269 (Z2013*) 1 0.83%
R1b TOTAL 14 11.67%
Haplogroup R1a — 9 (7.50%)
Subclade Count % of Total
R1a (unspecified) 8 6.67%
R-Z92 (R1a-Z280) 1 0.83%
R1a TOTAL 9 7.50%
Haplogroup T — 3 (2.50%)
Subclade Count % of Total
T1a 3 2.50%
Haplogroup G — 1 (0.83%)
Subclade Count % of Total
G2a-L497 1 0.83%
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SOUTHWEST REGION — Y-DNA Haplogroup Distribution
Total Sample Size (N = 180)
1. Macro-Haplogroup Summary
Haplogroup Count % of Total
J 59 32.78%
E (E1b1b) 38 21.11%
I (I1 + I2) 34 18.89%
R1b 22 12.22%
R1a 17 9.44%
G 5 2.78%
T 2 1.11%
L 2 1.11%
N 1 0.56%
TOTAL 180 100.00%
2. Detailed Haplogroup & Subclade Breakdown
Haplogroup E (E1b1b) — 38 (21.11%)
Subclade Count % of Total
E-V13 (unspecified) 25 13.89%
E-V13 → Z1057 3 1.67%
E-V13 → Z5017 4 2.22%
E-CTS1273 1 0.56%
E-CTS2001 1 0.56%
E-L241 1 0.56%
E-M123 1 0.56%
E-Y7168 1 0.56%
E-V37092 1 0.56%
E TOTAL 38 21.11%
Haplogroup J — 59 (32.78%)
J2a — 38 (21.11%)
Subclade Count % of Total
J2a (unspecified) 30 16.67%
J2a-L70 4 2.22%
J-M92 (J2a) 2 1.11%
J2a-L24 1 0.56%
J2a-CTS333 1 0.56%
J2a subtotal 38 21.11%
J2b — 11 (6.11%)
Subclade Count % of Total
J2b-Z631 5 2.78%
J2b-M241 2 1.11%
J2b-M205 → Y22059 1 0.56%
J2b2-M102 1 0.56%
J-Z1043 (J2b) 1 0.56%
J2b (unspecified) 1 0.56%
J2b subtotal 11 6.11%
J1 — 10 (5.56%)
Subclade Count % of Total
J1 (unspecified) 3 1.67%
J1-Z1828 1 0.56%
J1-P58 → Z2313 4 2.22%
J-CTS5368 (J1) 1 0.56%
J-Z2313 1 0.56%
J1 subtotal 10 5.56%
J TOTAL: 59 (32.78%)
Haplogroup I — 34 (18.89%)
I2 — 31 (17.22%)
Subclade Count % of Total
I2-Z17855 17 9.44%
I2-Y3120 6 3.33%
I-S17250 3 1.67%
I-CTS5966 3 1.67%
I2-CTS10228 1 0.56%
I-Z26381 1 0.56%
I2 subtotal 31 17.22%
I1 — 3 (1.67%)
Subclade Count % of Total
I1-P109 → FGC22045 1 0.56%
I1 (unspecified) 2 1.11%
I1 subtotal 3 1.67%
I TOTAL: 34 (18.89%)
Haplogroup R1b — 22 (12.22%)
Subclade Count % of Total
R1b-Z2013 17 9.45%
R-L23 3 1.67%
R1b-U152 → L2 1 0.56%
R1b-Z2103 → Z2705 1 0.56%
R1b TOTAL 22 12.22%
Haplogroup R1a — 17 (9.44%)
Subclade Count % of Total
R1a (unspecified) 9 5.00%
R1a-Z280 4 2.22%
R1a-M458 / YP417 / A11460 3 1.67%
R-Y57 (Z280) 1 0.56%
R1a TOTAL 17 9.44%
Haplogroup G — 5 (2.78%)
Subclade Count % of Total
G2a (all) 5 2.78%
Haplogroup T — 2 (1.11%)
Subclade Count % of Total
T-L446 2 1.11%
Haplogroup L — 2 (1.11%)
Subclade Count % of Total
L-M22 2 1.11%
Haplogroup N — 1 (0.56%)
Subclade Count % of Total
N-TAT → L1026 1 0.56%
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Southeast Region — Y-DNA Haplogroup Structure
Total Sample Size (n = 110)
1. Overall Haplogroup Distribution
Haplogroup Count % of Total
E (E1b1b) 39 35.45%
I (I2) 29 26.36%
J (J1 + J2a) 19 17.27%
R1b 13 11.82%
R1a 8 7.27%
G (G2a) 2 1.82%
TOTAL 110 100.00%
2. Detailed Haplogroup & Subclade Breakdown
Haplogroup E (E1b1b) — 39 (35.45%)
Subclade Count % of Total
E-V13 (unspecified) 31 28.18%
E-V13 → CTS5856 → Y16729 4 3.64%
E-V13 → CTS5856 (unspecified) 2 1.82%
E-V13 → Z1057 1 0.91%
E-L117 / E1b (unspecified) 1 0.91%
E TOTAL 39 35.45%
Haplogroup I (I2) — 29 (26.36%)
Subclade Count % of Total
I2a (unspecified) 24 21.82%
I2a-CTS10228 4 3.64%
I2-Dinaric North 1 0.91%
I TOTAL 29 26.36%
Haplogroup J — 19 (17.27%)
Subclade Count % of Total
J2a 16 14.55%
J1 3 2.73%
J TOTAL 19 17.27%
Haplogroup R1a — 8 (7.27%)
Subclade Count % of Total
R1a (unspecified) 8 7.27%
R1a TOTAL 8 7.27%
Haplogroup R1b — 13 (11.82%)
Subclade Count % of Total
R1b-Z2013 10 9.09%
R1b-L506 (incl. L278 / M415) 3 2.73%
R1b TOTAL 13 11.82%
Haplogroup G (G2a) — 2 (1.82%)
Subclade Count % of Total
G2a (incl. L30 → P303) 2 1.82%
G TOTAL 2 1.82%
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Macedonian Turks — Y-Chromosome Haplogroup Distribution
Total Sample Size (N = 109)
Overall Haplogroup Composition
Haplogroup / Subclade Count % of Total
J2a-M410 38 34.86%
R1b-M269 22 20.18%
I2-P215 21 19.27%
E1b-M35 13 11.93%
I1-M253 4 3.67%
J1-M267 2 1.83%
N-Tat 2 1.83%
R1a-M198 2 1.83%
I2a-M223 2 1.83%
J2b-M102 1 0.92%
R1b-U152 1 0.92%
KLT-M9 1 0.92%
TOTAL 109 100.00%
