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Patterson et al. 2022 study examined 18 samples from the Middle Bronze Age up to Early Iron Age Croatia, which was part of Illyria. Out of the nine Y-DNA samples retrieved, which coincide with the historical territory where Illyrians lived (including tested Iapydes and Liburni sites), almost all belonged to the patrilineal line J2b2a1-L283 (>J-PH1602 > J-Y86930 and >J-Z1297 subclades) with the exception of one R1b-L2. The mtDNA haplogroups fell under various subclades of H, H1, H3b, H5, J1c2, J1c3, T2a1a, T2b, T2b23, U5a1g, U8b1b1, HV0e. In a three-way admixture model, they approximately had 49-59% EEF, 35-46% Steppe and 2-10% WHG-related ancestry.[51] In Lazaridis et al. (2022) key parts of the territory of historical territory of Illyria were tested. In 18 samples from the Cetina culture, all males except for one (R-L51 > Z2118) carried Y-DNA haplogroup J-L283. Many of them could be further identified as J-L283 > Z597 (> J-Y15058 > J-Z38240 > J-PH1602). The majority of individuals carried mtDNA haplogroups J1c1 and H6a1a. The related Posušje culture yielded the same Y-DNA haplogroup (J-L283 > J-Z38240). The same J-L283 population appears in the MBA-IA Velim Kosa tumuli of Liburni in Croatia (J-PH1602), and similar in LBA-IA Velika Gruda tumuli in Montenegro (J-Z2507 > J-Z1297 > J-Y21878). The oldest J-L283 (> J-Z597) sample in the study was found in MBA Shkrel, northern Albania as early as the 19th century BCE. In northern Albania, IA Çinamak, half of them men carried J-L283 (> J-Z622, J-Y21878) and the other half R-M269 (R-CTS1450, R-PF7563). The oldest sample in Çinamak dates to the first era of post-Yamnaya movements (EBA) and carries R-M269.[52] Autosomally, Croatian Bronze Age samples from various sites, from Cetina valley and Bezdanjača Cave were "extremely similar in their ancestral makeup",[53] while from Montenegro's Velika Gruda mainly had an admixture of "Anatolian Neolithic (~50%), Eastern European hunter-gatherer (~12%), and Balkan hunter-gatherer ancestry (~18%)".[54] The oldest Balkan J-L283 samples have been found in final Early Bronze Age (ca. 1950 BCE) site of Mokrin in Serbia and about 100-150 years later in Shkrel, northern Albania.[55][56]
