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In Serbian tribes it's based on male founder as well (seed), however taking a foreign woman is still discouraged, it happens from time to time. I was curious about your codes of conducts to compare, what u call "Pashtunwali" for example in Serbian system,
If u descend from same man, u cannot marry anyone inside it. So even if u have to back a 1000 years to get a common ancestor, u still cannot marry. Usually villages are based on one or two tribes, and they cannot marry within the village.
Second, if u do not descend from same man, u have to trace ur ancestry to 8 generations to make sure there is no common link. For example if ur great-grandmother came from the same family (here clan, not tribe) u cannot marry even though this is all female lines.
I find the above two most important, as it underlines how people organize. For example, I still can go to someone from the same male founder, and we trace to 1300s, and he will treat me how an American will treat a first cousin, or even better. At same time, I keep in touch with only immediate cousins form my female line.