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Macedonian Y-DNA
Y-Chromosome Haplogroup Distribution – Vardar Region
Sample size (N) = 120
1. Major Haplogroup Composition
Haplogroup Count Percentage
E 36 30.00%
I 28 23.33%
J 24 20.00%
R1b 21 17.50%
R1a 10 8.33%
L 1 0.83%
TOTAL 120 100.00%
2. Detailed Haplogroup & Subclade Breakdown
Haplogroup E – 36 (30.00%)
E-V13 Cluster – 29 (24.17%)
Subclade Count Percentage
E-V13 (unspecified) 26 21.67%
E-V13 (regional / unresolved) 3 2.50%
E-V13 subtotal 29 24.17%
Other E Lineages – 7 (5.83%)
Subclade Count Percentage
E-V22 2 1.67%
E-L677 2 1.67%
E-CTS2548 1 0.83%
E-PH1173 1 0.83%
E-M10 (E-V68) 1 0.83%
Other E subtotal 7 5.83%
E TOTAL: 36 (30.00%)
Haplogroup I – 28 (23.33%)
I2 Lineages – 26 (21.67%)
Subclade Count Percentage
I2a (unspecified) 12 10.00%
I-CTS10057 (incl. CTS10100, Z162) 6 5.00%
I-CTS5966 4 3.33%
I2-Y3120 (incl. PH908, S17250) 4 3.33%
I2 subtotal 26 21.67%
Other I – 2 (1.67%)
Subclade Count Percentage
I (unspecified) 2 1.67%
I TOTAL: 28 (23.33%)
Haplogroup J – 24 (20.00%)
J2 Lineages – 21 (17.50%)
Subclade Count Percentage
J2a (unspecified) 13 10.83%
J2a-L24 1 0.83%
J-L70 (J2a) 1 0.83%
J-M67 (J2a) 3 2.50%
J2b-M241 / Z38299 3 2.50%
J2 subtotal 21 17.50%
J1 – 3 (2.50%)
Subclade Count Percentage
J1 3 2.50%
J TOTAL: 24 (20.00%)
Haplogroup R1b – 21 (17.50%)
R1b-Z2013 – 18 (15.00%)
Subclade Count Percentage
R1b-Z2013 18 15.00%
R1b-L421 Cluster – 3 (2.50%)
Subclade Count Percentage
R1b-L421 / downstream 3 2.50%
R1b TOTAL: 21 (17.50%)
Haplogroup R1a – 10 (8.33%)
Subclade Count Percentage
R1a (unspecified) 7 5.83%
R-Z93 (R1a) 1 0.83%
R1a (regional clusters) 2 1.67%
R1a TOTAL: 10 (8.33%)
Haplogroup L – 1 (0.83%)
Subclade Count Percentage
L-L1307 1 0.83%
L TOTAL: 1 (0.83%)
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SKOPJE REGION — Y-DNA Haplogroup Distribution
Total samples (N = 120)
1. Overall Haplogroup Frequencies
Haplogroup Count Percentage
I 33 27.50%
E 30 25.00%
J 24 20.00%
R1b 21 17.50%
R1a 10 8.33%
G 2 1.67%
TOTAL 120 100.00%
2. Haplogroup E — 30 (25.00%)
Subclade Count Percentage
E1b1b1a (xV13) 2 1.67%
E-V13 (unspecified) 23 19.17%
E-V13 → Z1057 3 2.50%
E-V13 → Z5018 1 0.83%
E-V13 → Z5017 1 0.83%
E TOTAL 30 25.00%
3. Haplogroup J — 24 (20.00%)
J2a — 14 (11.67%)
Subclade Count Percentage
J2a (unspecified) 4 3.33%
J-M67 / J2a1b / CTS6372 4 3.33%
J-L24 3 2.50%
J-M410 (xL24) 1 0.83%
J-M92 → PF7412 1 0.83%
J-L26 1 0.83%
J2a TOTAL 14 11.67%
J2b — 9 (7.50%)
Subclade Count Percentage
J2b (unspecified) 4 3.33%
J2b-L283 3 2.50%
J2b-M205 → Y22059 2 1.67%
J2b TOTAL 9 7.50%
J1 — 1 (0.83%)
Subclade Count Percentage
J1-M267 1 0.83%
J TOTAL: 24 (20.00%)
4. Haplogroup I — 33 (27.50%)
I1 — 3 (2.50%)
Subclade Count Percentage
I1 (unspecified) 1 0.83%
I1-P109 → FGC22045 2 1.67%
I1 TOTAL 3 2.50%
I2 — 30 (25.00%)
Subclade Count Percentage
I2a (unspecified) 3 2.50%
I2-CTS5966 1 0.83%
I2-CTS10228 3 2.50%
I2-Y3120 (total) 23 19.17%
I2-Y3120 Breakdown
Subclade Count Percentage
Y3120 (unspecified) 7 5.83%
S17250 (xPH908) 4 3.33%
S17250 → PH908 3 2.50%
Y4884 3 2.50%
Z17855 6 5.00%
Y3120 TOTAL 23 19.17%
I TOTAL: 33 (27.50%)
5. Haplogroup R1a — 10 (8.33%)
Subclade Count Percentage
R1a (unspecified) 6 5.00%
R1a-Z280 → Y2902 2 1.67%
R1a-M458 → L1029 2 1.67%
R1a TOTAL 10 8.33%
6. Haplogroup R1b — 21 (17.50%)
Subclade Count Percentage
R1b-M269 → Z2013 12 10.00%
R1b-L277 3 2.50%
R1b-L421 3 2.50%
R1b-L23 (xZ2103) 2 1.67%
R1b-M269 → L51 → P311 1 0.83%
R1b TOTAL 21 17.50%
7. Haplogroup G — 2 (1.67%)
Subclade Count Percentage
G2a-L497 → L42 1 0.83%
G2a-L497 → L42 → Y128028 1 0.83%
G TOTAL 2 1.67%
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POLOG REGION — Y-DNA Haplogroup Distribution
Total samples (N = 110)
1. Major Haplogroups
Haplogroup Count Percentage
J 34 30.91%
E 23 20.91%
I 23 20.91%
R1a 14 12.73%
R1b 13 11.82%
T 2 1.82%
G 1 0.91%
TOTAL 110 100.00%
2. Haplogroup J — 34 (30.91%)
J2a — 13 (11.82%)
Subclade Count Percentage
J2a-L70 (incl. L397, L398 and derived) 9 8.18%
J2a (unspecified) 4 3.64%
J2a TOTAL 13 11.82%
J2b — 20 (18.18%)
Subclade Count Percentage
J2b (unspecified) 9 8.18%
J2b-L283 (incl. FT134628, Z631) 8 7.27%
J2b-M241 3 2.73%
J2b TOTAL 20 18.18%
J1 — 1 (0.91%)
Subclade Count Percentage
J1-P58 / J1e 1 0.91%
3. Haplogroup E — 23 (20.91%)
Subclade Count Percentage
E-V13 (all downstream clades) 20 18.18%
E-CTS1273 (incl. CTS2374, CTS5371) 2 1.82%
E-L241 1 0.91%
E TOTAL 23 20.91%
4. Haplogroup I — 23 (20.91%)
Subclade Count Percentage
I2-Y3120 (all) 21 19.09%
└─ I2-S17250 → PH908 6 5.45%
I2c-L596 1 0.91%
I2a (unspecified) 1 0.91%
I TOTAL 23 20.91%
5. Haplogroup R1a — 14 (12.73%)
Subclade Count Percentage
R1a-M458 → L1029 2 1.82%
R1a-M417 / M512 8 7.27%
R1a (unspecified) 4 3.64%
R1a TOTAL 14 12.73%
6. Haplogroup R1b — 13 (11.82%)
Subclade Count Percentage
R1b-Z2013 (incl. Y32147) 6 5.45%
R1b-CTS9219 2 1.82%
R1b-P312 1 0.91%
R1b-Z2103 → Z2705 2 1.82%
R1b-PF7558 / PF7562 2 1.82%
R1b TOTAL 13 11.82%
7. Minor Haplogroups
Haplogroup Count Percentage
T-Y11151 2 1.82%
G2a-M406 1 0.91%
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NORTHEAST REGION — Y-DNA Haplogroup Distribution
Total samples (N = 110)
1. Major Haplogroups
Haplogroup Count Percentage
I 34 30.91%
E 28 25.45%
R1b 20 18.18%
J 15 13.64%
R1a 8 7.27%
G 5 4.55%
TOTAL 110 100.00%
2. Haplogroup I — 34 (30.91%)
Subclade Count Percentage
I2-Y3120 (all downstream) 26 23.64%
├─ I2-S17250 → PH908 11 10.00%
I2-CTS11030 1 0.91%
I2-CTS595 1 0.91%
I2-M223 → P78 → Y4884 2 1.82%
I2-S17250 (undifferentiated) 2 1.82%
I2a (unspecified) 2 1.82%
I TOTAL 34 30.91%
3. Haplogroup E — 28 (25.45%)
Subclade Count Percentage
E-V13 (all downstream clades) 22 20.00%
├─ Z1057 / Z5017 / Z5018 (included above) —
E-L241 2 1.82%
E-L250 2 1.82%
E-BY5610 1 0.91%
E-M123 1 0.91%
E TOTAL 28 25.45%
4. Haplogroup R1b — 20 (18.18%)
Subclade Count Percentage
R1b-Z2013 (incl. FT330613) 11 10.00%
R1b-L277 2 1.82%
R1b-Z2103 → Z2705 / Y32147 3 2.73%
R1b-U152 1 0.91%
R1b-R247 2 1.82%
R1b (unspecified M269) 1 0.91%
R1b TOTAL 20 18.18%
5. Haplogroup J — 15 (13.64%)
Subclade Count Percentage
J2b-M205 → Y22059 (incl. Y22063) 11 10.00%
J2a (incl. L70) 2 1.82%
J1-P58 → Z2313 1 0.91%
J2b (unspecified) 1 0.91%
J TOTAL 15 13.64%
6. Haplogroup R1a — 8 (7.27%)
Subclade Count Percentage
R1a-Z280 (incl. YP4278, YP278.2) 4 3.64%
R1a (unspecified) 4 3.64%
R1a TOTAL 8 7.27%
7. Haplogroup G — 5 (4.55%)
Subclade Count Percentage
G2a3 2 1.82%
G2a3b1 1 0.91%
G2a-PF3359 1 0.91%
G-Z2022 1 0.91%
G TOTAL 5 4.55%
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Y-Chromosome Haplogroup Distribution – East Region
Sample size (N = 110)
1. Major Haplogroup Composition
Haplogroup Count Percentage
I 34 30.91%
E 31 28.18%
R1b 22 20.00%
J 16 14.55%
R1a 7 6.36%
TOTAL 110 100.00%
2. Detailed Haplogroup and Subclade Structure
Haplogroup I — 34 (30.91%)
I2 Lineages — 32 (29.09%)
Subclade Count Percentage
I2a (unspecified) 6 5.45%
I2-CTS10228 (incl. CTS5966) 10 9.09%
I2-M423 8 7.27%
I-L68 (I2) 7 6.36%
I2 subtotal 31 28.18%
I1 Lineages — 2 (1.82%)
Subclade Count Percentage
I1-P109 2 1.82%
Haplogroup I TOTAL: 34 (30.91%)
Haplogroup E — 31 (28.18%)
E-V13 Cluster — 29 (26.36%)
Subclade Count Percentage
E-V13 (unspecified) 24 21.82%
E-CTS1273 (CTS2374 / CTS5371) 4 3.64%
E-V13 → CTS5856 → Z5018 → S2979 → FGC11457 1 0.91%
E-V13 subtotal 29 26.36%
Other E Lineages — 2 (1.82%)
Subclade Count Percentage
E (unresolved / non-V13) 2 1.82%
Haplogroup E TOTAL: 31 (28.18%)
Haplogroup R1b — 22 (20.00%)
R1b-Z2103 (Balkan–Carpathian Cluster) — 3 (2.73%)
Subclade Count Percentage
R1b-M269 → L23 → Z2103 → L584 3 2.73%
R1b-Z2013 — 13 (11.81%)
Subclade Count Percentage
R1b-Z2013 13 11.81%
Other R1b Lineages — 6 (5.45%)
Subclade Count Percentage
R1b-Z156 2 1.82%
R-PF7558 (R1b) 1 0.91%
R1b (unspecified) 3 2.73%
Haplogroup R1b TOTAL: 22 (20.00%)
Haplogroup J — 16 (14.55%)
J2 Lineages — 14 (12.73%)
Subclade Count Percentage
J2a-L26 6 5.45%
J-M67 / J2a1b (CTS6372 / PF5120) 3 2.73%
J2a (unspecified) 5 4.55%
Other J Lineages — 2 (1.82%)
Subclade Count Percentage
J2b 1 0.91%
J1-CTS5368 1 0.91%
Haplogroup J TOTAL: 16 (14.55%)
Haplogroup R1a — 7 (6.36%)
Subclade Count Percentage
R1a-L1029 4 3.64%
R1a (unspecified) 3 2.73%
R1a TOTAL 7 6.36%
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Pelagonia Region — Y-Chromosome Haplogroup Distribution
Total Sample Size (N = 120)
1. Major Haplogroup Composition
Haplogroup Count % of Total
E 39 32.50%
I 27 22.50%
J 27 22.50%
R1b 14 11.67%
R1a 9 7.50%
T 3 2.50%
G 1 0.83%
TOTAL 120 100.00%
2. Detailed Haplogroup & Subclade Breakdown
Haplogroup E — 39 (32.50%)
E-V13 Cluster — 33 (27.50%)
Subclade Count % of Total
E-V13 (unspecified) 30 25.00%
E-V13 → CTS5856 → S7461 1 0.83%
E-V13 → CTS5856 → Z5018 → S2979 → Y3183 1 0.83%
E-V13 → CTS5856 → Z5017 → Z5016 → CTS9320 / S26015 1 0.83%
E-V13 subtotal 33 27.50%
Other E Lineages — 6 (5.00%)
Subclade Count % of Total
E-CTS1273 4 3.33%
E-Z1057 2 1.67%
E TOTAL: 39 (32.50%)
Haplogroup I — 27 (22.50%)
I2 Lineages — 23 (19.17%)
Subclade Count % of Total
I2a (unspecified) 9 7.50%
I-M423 6 5.00%
I-S17250 5 4.17%
I-CTS10057 3 2.50%
I2 subtotal 23 19.17%
Other I — 4 (3.33%)
Subclade Count % of Total
I1-L759 2 1.67%
I-Z17855 1 0.83%
I-P78 1 0.83%
I TOTAL: 27 (22.50%)
Haplogroup J — 27 (22.50%)
J2 Lineages — 22 (18.33%)
Subclade Count % of Total
J2a (unspecified) 15 12.50%
J2a-L70 1 0.83%
J2b-L283 4 3.33%
J-M241 (J2b) 1 0.83%
J-FT278595 (J2b) 1 0.83%
J2 subtotal 22 18.33%
J1 — 5 (4.17%)
Subclade Count % of Total
J1 (unspecified) 5 4.17%
J TOTAL: 27 (22.50%)
Haplogroup R1b — 14 (11.67%)
Subclade Count % of Total
R1b-Z2013 11 9.17%
R1b-Z156 1 0.83%
R1b1b2a 1 0.83%
R1b-M269 (Z2013*) 1 0.83%
R1b TOTAL 14 11.67%
Haplogroup R1a — 9 (7.50%)
Subclade Count % of Total
R1a (unspecified) 8 6.67%
R-Z92 (R1a-Z280) 1 0.83%
R1a TOTAL 9 7.50%
Haplogroup T — 3 (2.50%)
Subclade Count % of Total
T1a 3 2.50%
Haplogroup G — 1 (0.83%)
Subclade Count % of Total
G2a-L497 1 0.83%
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SOUTHWEST REGION — Y-DNA Haplogroup Distribution
Total Sample Size (N = 180)
1. Macro-Haplogroup Summary
Haplogroup Count % of Total
J 59 32.78%
E (E1b1b) 38 21.11%
I (I1 + I2) 34 18.89%
R1b 22 12.22%
R1a 17 9.44%
G 5 2.78%
T 2 1.11%
L 2 1.11%
N 1 0.56%
TOTAL 180 100.00%
2. Detailed Haplogroup & Subclade Breakdown
Haplogroup E (E1b1b) — 38 (21.11%)
Subclade Count % of Total
E-V13 (unspecified) 25 13.89%
E-V13 → Z1057 3 1.67%
E-V13 → Z5017 4 2.22%
E-CTS1273 1 0.56%
E-CTS2001 1 0.56%
E-L241 1 0.56%
E-M123 1 0.56%
E-Y7168 1 0.56%
E-V37092 1 0.56%
E TOTAL 38 21.11%
Haplogroup J — 59 (32.78%)
J2a — 38 (21.11%)
Subclade Count % of Total
J2a (unspecified) 30 16.67%
J2a-L70 4 2.22%
J-M92 (J2a) 2 1.11%
J2a-L24 1 0.56%
J2a-CTS333 1 0.56%
J2a subtotal 38 21.11%
J2b — 11 (6.11%)
Subclade Count % of Total
J2b-Z631 5 2.78%
J2b-M241 2 1.11%
J2b-M205 → Y22059 1 0.56%
J2b2-M102 1 0.56%
J-Z1043 (J2b) 1 0.56%
J2b (unspecified) 1 0.56%
J2b subtotal 11 6.11%
J1 — 10 (5.56%)
Subclade Count % of Total
J1 (unspecified) 3 1.67%
J1-Z1828 1 0.56%
J1-P58 → Z2313 4 2.22%
J-CTS5368 (J1) 1 0.56%
J-Z2313 1 0.56%
J1 subtotal 10 5.56%
J TOTAL: 59 (32.78%)
Haplogroup I — 34 (18.89%)
I2 — 31 (17.22%)
Subclade Count % of Total
I2-Z17855 17 9.44%
I2-Y3120 6 3.33%
I-S17250 3 1.67%
I-CTS5966 3 1.67%
I2-CTS10228 1 0.56%
I-Z26381 1 0.56%
I2 subtotal 31 17.22%
I1 — 3 (1.67%)
Subclade Count % of Total
I1-P109 → FGC22045 1 0.56%
I1 (unspecified) 2 1.11%
I1 subtotal 3 1.67%
I TOTAL: 34 (18.89%)
Haplogroup R1b — 22 (12.22%)
Subclade Count % of Total
R1b-Z2013 17 9.45%
R-L23 3 1.67%
R1b-U152 → L2 1 0.56%
R1b-Z2103 → Z2705 1 0.56%
R1b TOTAL 22 12.22%
Haplogroup R1a — 17 (9.44%)
Subclade Count % of Total
R1a (unspecified) 9 5.00%
R1a-Z280 4 2.22%
R1a-M458 / YP417 / A11460 3 1.67%
R-Y57 (Z280) 1 0.56%
R1a TOTAL 17 9.44%
Haplogroup G — 5 (2.78%)
Subclade Count % of Total
G2a (all) 5 2.78%
Haplogroup T — 2 (1.11%)
Subclade Count % of Total
T-L446 2 1.11%
Haplogroup L — 2 (1.11%)
Subclade Count % of Total
L-M22 2 1.11%
Haplogroup N — 1 (0.56%)
Subclade Count % of Total
N-TAT → L1026 1 0.56%
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Southeast Region — Y-DNA Haplogroup Structure
Total Sample Size (n = 110)
1. Overall Haplogroup Distribution
Haplogroup Count % of Total
E (E1b1b) 39 35.45%
I (I2) 29 26.36%
J (J1 + J2a) 19 17.27%
R1b 13 11.82%
R1a 8 7.27%
G (G2a) 2 1.82%
TOTAL 110 100.00%
2. Detailed Haplogroup & Subclade Breakdown
Haplogroup E (E1b1b) — 39 (35.45%)
Subclade Count % of Total
E-V13 (unspecified) 31 28.18%
E-V13 → CTS5856 → Y16729 4 3.64%
E-V13 → CTS5856 (unspecified) 2 1.82%
E-V13 → Z1057 1 0.91%
E-L117 / E1b (unspecified) 1 0.91%
E TOTAL 39 35.45%
Haplogroup I (I2) — 29 (26.36%)
Subclade Count % of Total
I2a (unspecified) 24 21.82%
I2a-CTS10228 4 3.64%
I2-Dinaric North 1 0.91%
I TOTAL 29 26.36%
Haplogroup J — 19 (17.27%)
Subclade Count % of Total
J2a 16 14.55%
J1 3 2.73%
J TOTAL 19 17.27%
Haplogroup R1a — 8 (7.27%)
Subclade Count % of Total
R1a (unspecified) 8 7.27%
R1a TOTAL 8 7.27%
Haplogroup R1b — 13 (11.82%)
Subclade Count % of Total
R1b-Z2013 10 9.09%
R1b-L506 (incl. L278 / M415) 3 2.73%
R1b TOTAL 13 11.82%
Haplogroup G (G2a) — 2 (1.82%)
Subclade Count % of Total
G2a (incl. L30 → P303) 2 1.82%
G TOTAL 2 1.82%
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Macedonian Turks — Y-Chromosome Haplogroup Distribution
Total Sample Size (N = 109)
Overall Haplogroup Composition
Haplogroup / Subclade Count % of Total
J2a-M410 38 34.86%
R1b-M269 22 20.18%
I2-P215 21 19.27%
E1b-M35 13 11.93%
I1-M253 4 3.67%
J1-M267 2 1.83%
N-Tat 2 1.83%
R1a-M198 2 1.83%
I2a-M223 2 1.83%
J2b-M102 1 0.92%
R1b-U152 1 0.92%
KLT-M9 1 0.92%
TOTAL 109 100.00%
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Macedonian Aromanians: Y-Haplogroup Distribution (N=108)
Haplogroup Kruševo (n=43) Štip (n=65) Total Count Combined Percentage
R1b 12 15 27 25.0%
E1b1b 9 12 21 19.4%
I 9 11 20 18.5%
R1a 5 14 19 17.6%
J 5 13 18 16.7%
G 3 0 3 2.8%
Total 43 65 108 100%
Overall Haplogroup Distribution
Haplogroup Štip (n=65) Kruševo (n=43) Total (n=108)
C-V20 0 (0.00%) 0 (0.00%) 0 (0.00%)
E-M81 0 (0.00%) 1 (2.33%) 1 (0.93%)
E-V22 0 (0.00%) 2 (4.65%) 2 (1.85%)
E-V13 12 (18.46%) 6 (13.95%) 18 (16.67%)
G1 0 (0.00%) 0 (0.00%) 0 (0.00%)
G2a 0 (0.00%) 3 (6.98%) 3 (2.78%)
I1 2 (3.08%) 1 (2.33%) 3 (2.78%)
I2-Y196 0 (0.00%) 8 (18.60%) 8 (7.41%)
I2-M223 3 (4.62%) 0 (0.00%) 3 (2.78%)
I2 (unspecified) 6 (9.23%) 0 (0.00%) 6 (5.56%)
J1 0 (0.00%) 0 (0.00%) 0 (0.00%)
J2a 13 (20.00%) 3 (6.98%) 16 (14.81%)
J2b-M205 0 (0.00%) 0 (0.00%) 0 (0.00%)
J2b-M241 0 (0.00%) 2 (4.65%) 2 (1.85%)
T 0 (0.00%) 0 (0.00%) 0 (0.00%)
N1c 0 (0.00%) 0 (0.00%) 0 (0.00%)
R1a 14 (21.54%) 5 (11.63%) 19 (17.59%)
R1b (Western) 0 (0.00%) 4 (9.30%) 4 (3.70%)
R1b (Eastern) 15 (23.08%) 8 (18.60%) 23 (21.30%)
TOTAL 65 (100.00%) 43 (100.00%) 108 (100.00%)
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TOTAL SAMPLE SIZE
N = 980
1. Overall Haplogroup Composition
Haplogroup Count %
E (E1b1b) 264 26.94
I 242 24.69
J 218 22.24
R1b 146 14.90
R1a 83 8.47
G 16 1.63
T 7 0.71
L 3 0.31
N 1 0.10
Total 980 100.00
2. Haplogroup E (E1b1b) — 264 (26.94%)
2.1 E-V13 — 225 (22.96%)
Subclade Count %
E-V13 (unspecified / unresolved) 204 20.82
E-CTS1273 (V13) 7 0.71
E-V13 > Z1057 4 0.41
E-V13 > Z5017 4 0.41
E-V13 > CTS5856 (unspecified) 2 0.20
E-V13 > CTS5856 > Y16729 4 0.41
E-V13 Total 225 22.96
2.2 Non-V13 E Lineages — 39 (3.98%)
Subclade Count %
E-CTS1273 (non-V13) 4 0.41
E-L241 4 0.41
E-Z1057 (non-V13) 2 0.20
E-L250 2 0.20
E-V22 2 0.20
E-L677 2 0.20
E-M123 2 0.20
E-L117 / E1b (unspecified) 1 0.10
E-CTS2001 1 0.10
E-Y7168 1 0.10
E-V37092 1 0.10
E-CTS2548 1 0.10
E-PH1173 1 0.10
E-M10 (V68) 1 0.10
E (unresolved / other) 14 1.43
Non-V13 Total 39 3.98
3. Haplogroup I — 242 (24.69%)
3.1 I1 — 10 (1.02%)
Subclade Count %
I1-P109 (all downstream) 6 0.61
I1-L759 2 0.20
I1 (unspecified) 2 0.20
I1 Total 10 1.02
3.2 I2 — 232 (23.67%)
Subclade Count %
I2-Y3120 (all downstream) 110 11.22
I2-CTS10228 (incl. CTS5966) 25 2.55
I2-M423 (xY3120) 14 1.43
I2-Dinaric North 1 0.10
I-L68 (I2) 7 0.71
I2a (unspecified) 59 6.02
I2-CTS10057 (incl. CTS10100, Z162) 9 0.92
I2 (other / unspecified) 7 0.71
I2 Total 232 23.67
3.3 I2-Y3120 Breakdown — 110 (11.22%)
Subclade Count %
PH908 22 2.24
Y4884 7 0.71
Z17855 24 2.45
S17250 (undifferentiated) 16 1.63
Y3120 (unspecified downstream) 41 4.18
Y3120 Total 110 11.22
4. Haplogroup J — 218 (22.24%)
4.1 J2a — 126 (12.86%)
Subclade Count %
J2a (unspecified) 89 9.08
J2a-L70 (incl. L397 / L398) 17 1.73
J-M67 / CTS6372 / PF5120 10 1.02
J-M92 2 0.20
J2a-L26 / L24 7 0.71
J2a-CTS333 1 0.10
J2a Total 126 12.86
4.2 J2b — 63 (6.43%)
Subclade Count %
J2b-L283 15 1.53
J2b-Z631 5 0.51
J2b-M205 (incl. Y22059 / Y22063) 14 1.43
J2b-M241 / Z38299 9 0.92
J2b2-M102 1 0.10
J-Z1043 (J2b) 1 0.10
J-FT278595 1 0.10
J2b (unspecified) 17 1.73
J2b Total 63 6.43
4.3 J1 — 29 (2.96%)
Subclade Count %
J1 (unspecified) 19 1.94
J1-P58 > Z2313 6 0.61
J1-CTS5368 2 0.20
J1-M267 1 0.10
J1-Z1828 1 0.10
J1 Total 29 2.96
5. Haplogroup R1b — 146 (14.90%)
Subclade Count %
R1b-Z2013 98 10.00
R-L23 3 0.31
R1b-Z2103 (incl. Z2705) 9 0.92
R1b-L421 6 0.61
R1b-L277 5 0.51
R1b-Z156 3 0.31
R1b-PF7558 / PF7562 3 0.31
R1b-CTS9219 2 0.20
R1b-R247 2 0.20
R1b-P312 / P311 2 0.20
R1b-U152 (incl. L2) 2 0.20
R1b-L506 (incl. L278 / M415) 3 0.31
R1b (unspecified M269) 8 0.82
R1b Total 146 14.90
6. Haplogroup R1a — 83 (8.47%)
Subclade Count %
R1a (unspecified) 49 5.00
R1a-Z280 (incl. Y57) 12 1.22
R1a-M458 → L1029 / YP417 11 1.12
R1a-M417 / M512 8 0.82
R-Z93 1 0.10
R1a (regional clusters) 2 0.20
R1a Total 83 8.47
7. Minor Haplogroups
G — 16 (1.63%)
Subclade Count %
G2a (all) 7 0.71
G2a-L497 / L42 3 0.31
G2a-M406 1 0.10
G2a3 2 0.20
G2a3b1 1 0.10
G2a-PF3359 1 0.10
G-Z2022 1 0.10
G Total 16 1.63
T — 7 (0.71%)
Subclade Count %
T-Y11151 2 0.20
T-L446 2 0.20
T1a 3 0.31
T Total 7 0.71
L — 3 (0.31%)
Subclade Count %
L-L1307 1 0.10
L-M22 2 0.20
L Total 3 0.31
N — 1 (0.10%)
Subclade Count %
N-TAT > L1026 1 0.10
N Total 1 0.10
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Y-DNA Haplogroup Distribution
Aromanians
Regions: Romania; Macedonia (Štip, Kruševo); Albania (Dukasi, Andon Poci)
Total samples: N = 214
1. Sample Breakdown by Location
Population Sample Size
Romania 48
Štip (Macedonia) 65
Kruševo (Macedonia) 43
Dukasi (Albania) 39
Andon Poci (Albania) 19
Total 214
2. Haplogroup Distribution by Population
Values shown as: Count (% within population)
Haplogroup Romania (n=48) Štip (n=65) Kruševo (n=43) Dukasi (n=39) Andon Poci (n=19) Total (n=214)
C-V20 2 (4.17%) 0 (0.00%) 0 (0.00%) 0 (0.00%) 0 (0.00%) 2 (0.93%)
E-M81 0 (0.00%) 0 (0.00%) 1 (2.33%) 0 (0.00%) 0 (0.00%) 1 (0.47%)
E-V22 0 (0.00%) 0 (0.00%) 2 (4.65%) 0 (0.00%) 0 (0.00%) 2 (0.93%)
E-V13 3 (6.25%) 12 (18.46%) 6 (13.95%) 7 (17.95%) 3 (15.79%) 31 (14.49%)
G1 0 (0.00%) 0 (0.00%) 0 (0.00%) 4 (10.26%) 0 (0.00%) 4 (1.87%)
G2a 0 (0.00%) 0 (0.00%) 3 (6.98%) 0 (0.00%) 0 (0.00%) 3 (1.40%)
I1 2 (4.17%) 2 (3.08%) 1 (2.33%) 2 (5.13%) 0 (0.00%) 7 (3.27%)
I2-Y196 10 (20.83%) 0 (0.00%) 8 (18.60%) 5 (12.82%) 8 (42.11%) 31 (14.49%)
I2-M223 2 (4.17%) 3 (4.62%) 0 (0.00%) 0 (0.00%) 0 (0.00%) 5 (2.34%)
I2 (unspecified) 0 (0.00%) 6 (9.23%) 0 (0.00%) 0 (0.00%) 0 (0.00%) 6 (2.80%)
J1 0 (0.00%) 0 (0.00%) 0 (0.00%) 1 (2.56%) 0 (0.00%) 1 (0.47%)
J2a 3 (6.25%) 13 (20.00%) 3 (6.98%) 0 (0.00%) 0 (0.00%) 19 (8.88%)
J2b-M205 0 (0.00%) 0 (0.00%) 0 (0.00%) 0 (0.00%) 1 (5.26%) 1 (0.47%)
J2b-M241 11 (22.92%) 0 (0.00%) 2 (4.65%) 18 (46.15%) 0 (0.00%) 31 (14.49%)
T 2 (4.17%) 0 (0.00%) 0 (0.00%) 0 (0.00%) 0 (0.00%) 2 (0.93%)
N1c 2 (4.17%) 0 (0.00%) 0 (0.00%) 0 (0.00%) 0 (0.00%) 2 (0.93%)
R1a 2 (4.17%) 14 (21.54%) 5 (11.63%) 1 (2.56%) 0 (0.00%) 22 (10.28%)
R1b (Western) 2 (4.17%) 0 (0.00%) 4 (9.30%) 1 (2.56%) 0 (0.00%) 7 (3.27%)
R1b (Eastern) 7 (14.58%) 15 (23.08%) 8 (18.60%) 0 (0.00%) 7 (36.84%) 37 (17.29%)
TOTAL 48 (100%) 65 (100%) 43 (100%) 39 (100%) 19 (100%) 214 (100%)
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NORTHEAST REGION
Y-DNA Haplogroup Distribution (N = 120)
1. Major Haplogroups Overview
Haplogroup Count Percentage
I 39 32.50%
E 30 25.00%
R1b 21 17.50%
J 15 12.50%
R1a 10 8.33%
G 5 4.17%
Total 120 100.00%
2. Haplogroup I
39 samples (32.50%)
Subclade Count Percentage
I2-Y3120 (all downstream) 26 21.67%
├─ I2-S17250 → PH908 11 9.17%
I2-CTS11030 1 0.83%
I2-CTS595 1 0.83%
I2-M223 → P78 → Y4884 2 1.67%
I2-S17250 (undifferentiated) 2 1.67%
I2a (unspecified) 7 5.83%
I total 39 32.50%
3. Haplogroup E
30 samples (25.00%)
Subclade Count Percentage
E-V13 (all downstream clades) 24 20.00%
├─ Z1057 / Z5017 / Z5018 (included above)
E-L241 2 1.67%
E-L250 2 1.67%
E-BY5610 1 0.83%
E-M123 1 0.83%
E total 30 25.00%
4. Haplogroup R1b
21 samples (17.50%)
Subclade Count Percentage
R1b-Z2013 (incl. FT330613) 12 10.00%
R1b-L277 2 1.67%
R1b-Z2103 → Z2705 / Y32147 3 2.50%
R1b-U152 1 0.83%
R1b-R247 2 1.67%
R1b (unspecified M269) 1 0.83%
R1b total 21 17.50%
5. Haplogroup J
15 samples (12.50%)
Subclade Count Percentage
J2b-M205 → Y22059 (incl. Y22063) 11 9.17%
J2a (incl. L70) 2 1.67%
J1-P58 → Z2313 1 0.83%
J2b (unspecified) 1 0.83%
J total 15 12.50%
6. Haplogroup R1a
10 samples (8.33%)
Subclade Count Percentage
R1a-Z280 (incl. YP4278, YP278.2) 4 3.33%
R1a (unspecified) 6 5.00%
R1a total 10 8.33%
7. Haplogroup G
5 samples (4.17%)
Subclade Count Percentage
G2a3 2 1.67%
G2a3b1 1 0.83%
G2a-PF3359 1 0.83%
G-Z2022 1 0.83%
G total 5 4.17%
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Y-Chromosome Haplogroup Distribution – East Region
Sample Size: N = 120
1. Major Haplogroup Composition
Haplogroup Count %
I 39 32.50
E 32 26.67
R1b 23 19.17
J 17 14.17
R1a 9 7.50
TOTAL 120 100.00
2. Detailed Haplogroup and Subclade Structure
HAPLOGROUP I — 39 (32.50%)
I2 Lineages — 37 (30.83%)
Subclade Count %
I2a (unspecified) 11 9.17
I2-CTS10228 (incl. CTS5966) 10 8.33
I2-M423 8 6.67
I-L68 (I2) 7 5.83
I2 (other / unspecified) 1 0.83
I2 Subtotal 37 30.83
I1 Lineages — 2 (1.67%)
Subclade Count %
I1-P109 2 1.67
Haplogroup I Total: 39 (32.50%)
HAPLOGROUP E — 32 (26.67%)
E-V13 Cluster — 30 (25.00%)
Subclade Count %
E-V13 (unspecified) 25 20.83
E-CTS1273 (CTS2374 / CTS5371) 4 3.33
E-V13 → CTS5856 → Z5018 → S2979 → FGC11457 1 0.83
E-V13 Subtotal 30 25.00
Other E Lineages — 2 (1.67%)
Subclade Count %
E (unresolved / non-V13) 2 1.67
Haplogroup E Total: 32 (26.67%)
HAPLOGROUP R1b — 23 (19.17%)
R1b-Z2103 (Balkan–Carpathian Cluster) — 3 (2.50%)
Subclade Count %
R1b-M269 → L23 → Z2103 → L584 3 2.50
R1b-Z2013 — 14 (11.67%)
Subclade Count %
R1b-Z2013 14 11.67
Other R1b Lineages — 6 (5.00%)
Subclade Count %
R1b-Z156 2 1.67
R-PF7558 (R1b) 1 0.83
R1b (unspecified) 3 2.50
Haplogroup R1b Total: 23 (19.17%)
HAPLOGROUP J — 17 (14.17%)
J2 Lineages — 16 (13.33%)
Subclade Count %
J2a-L26 6 5.00
J-M67 / J2a1b (CTS6372 / PF5120) 3 2.50
J2a (unspecified) 6 5.00
J2b 1 0.83
J2 Subtotal 16 13.33
J1 Lineages — 1 (0.83%)
Subclade Count %
J1-CTS5368 1 0.83
Haplogroup J Total: 17 (14.17%)
HAPLOGROUP R1a — 9 (7.50%)
Subclade Count %
R1a-L1029 4 3.33
R1a (unspecified) 5 4.17
Haplogroup R1a Total: 9 (7.50%)
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Y-DNA Haplogroup Analysis
Total Sample Size: N = 1000
OVERALL HAPLOGROUP COMPOSITION
Haplogroup Count %
E (E1b1b) 267 26.70
I 252 25.20
J 219 21.90
R1b 148 14.80
R1a 87 8.70
G 16 1.60
T 7 0.70
L 3 0.30
N 1 0.10
TOTAL 1000 100.00
HAPLOGROUP E (E1b1b) — 267 (26.70%)
E-V13 — 228 (22.80%)
Subclade Count %
E-V13 (unspecified / unresolved) 207 20.70
E-CTS1273 (V13) 7 0.70
E-V13 > Z1057 4 0.40
E-V13 > Z5017 4 0.40
E-V13 > CTS5856 (unspecified) 2 0.20
E-V13 > CTS5856 > Y16729 4 0.40
E-V13 Total 228 22.80
Non-V13 E Lineages — 39 (3.90%)
Subclade Count %
E-CTS1273 (non-V13) 4 0.40
E-L241 4 0.40
E-Z1057 (non-V13) 2 0.20
E-L250 2 0.20
E-V22 2 0.20
E-L677 2 0.20
E-M123 2 0.20
E-L117 / E1b (unspecified) 1 0.10
E-CTS2001 1 0.10
E-Y7168 1 0.10
E-V37092 1 0.10
E-CTS2548 1 0.10
E-PH1173 1 0.10
E-M10 (V68) 1 0.10
E (unresolved / other) 14 1.40
Non-V13 Total 39 3.90
HAPLOGROUP I — 252 (25.20%)
I1 — 10 (1.00%)
Subclade Count %
I1-P109 (all downstream) 6 0.60
I1-L759 2 0.20
I1 (unspecified) 2 0.20
I1 Total 10 1.00
I2 — 242 (24.20%)
Subclade Count %
I2-Y3120 (all downstream) 110 11.00
I2-CTS10228 (incl. CTS5966) 25 2.50
I2-M423 (xY3120) 14 1.40
I2-Dinaric North 1 0.10
I-L68 (I2) 7 0.70
I2a (unspecified) 69 6.90
I2-CTS10057 (incl. CTS10100, Z162) 9 0.90
I2 (other / unspecified) 7 0.70
I2 Total 242 24.20
I2-Y3120 Breakdown — 110 (11.00%)
Subclade Count %
PH908 22 2.20
Y4884 7 0.70
Z17855 24 2.40
S17250 (undifferentiated) 16 1.60
Y3120 (unspecified downstream) 41 4.10
Y3120 Total 110 11.00
HAPLOGROUP J — 219 (21.90%)
J2a — 127 (12.70%)
Subclade Count %
J2a (unspecified) 90 9.00
J2a-L70 (incl. L397 / L398) 17 1.70
J-M67 / CTS6372 / PF5120 10 1.00
J-M92 2 0.20
J2a-L26 / L24 7 0.70
J2a-CTS333 1 0.10
J2a Total 127 12.70
J2b — 63 (6.30%)
Subclade Count %
J2b-L283 15 1.50
J2b-Z631 5 0.50
J2b-M205 (incl. Y22059 / Y22063) 14 1.40
J2b-M241 / Z38299 9 0.90
J2b2-M102 1 0.10
J-Z1043 (J2b) 1 0.10
J-FT278595 1 0.10
J2b (unspecified) 17 1.70
J2b Total 63 6.30
J1 — 29 (2.90%)
Subclade Count %
J1 (unspecified) 19 1.90
J1-P58 > Z2313 6 0.60
J1-CTS5368 2 0.20
J1-M267 1 0.10
J1-Z1828 1 0.10
J1 Total 29 2.90
HAPLOGROUP R1b — 148 (14.80%)
Subclade Count %
R1b-Z2013 100 10.00
R-L23 3 0.30
R1b-Z2103 (incl. Z2705) 9 0.90
R1b-L421 6 0.60
R1b-L277 5 0.50
R1b-Z156 3 0.30
R1b-PF7558 / PF7562 3 0.30
R1b-CTS9219 2 0.20
R1b-R247 2 0.20
R1b-P312 / P311 2 0.20
R1b-U152 (incl. L2) 2 0.20
R1b-L506 (incl. L278 / M415) 3 0.30
R1b (unspecified M269) 8 0.80
R1b Total 148 14.80
HAPLOGROUP R1a — 87 (8.70%)
Subclade Count %
R1a (unspecified) 53 5.30
R1a-Z280 (incl. Y57) 12 1.20
R1a-M458 → L1029 / YP417 11 1.10
R1a-M417 / M512 8 0.80
R-Z93 1 0.10
R1a (regional clusters) 2 0.20
R1a Total 87 8.70
HAPLOGROUP G — 16 (1.60%)
Subclade Count %
G2a (all) 7 0.70
G2a-L497 / L42 3 0.30
G2a-M406 1 0.10
G2a3 2 0.20
G2a3b1 1 0.10
G2a-PF3359 1 0.10
G-Z2022 1 0.10
G Total 16 1.60
HAPLOGROUP T — 7 (0.70%)
Subclade Count %
T-Y11151 2 0.20
T-L446 2 0.20
T1a 3 0.30
T Total 7 0.70
HAPLOGROUP L — 3 (0.30%)
Subclade Count %
L-L1307 1 0.10
L-M22 2 0.20
L Total 3 0.30
HAPLOGROUP N — 1 (0.10%)
Subclade Count %
N-TAT > L1026 1 0.10
N Total 1 0.10
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