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''When we use DATES (19) to date the admixture of steppe ancestry in populations of Southeastern Europe (Fig. 5F and fig. S6), we arrive
at an estimate that this took place ~4850 years ago, i.e., precisely after the Yamnaya expansion, and within the time frame of our “highsteppe” cluster individuals. This suggests that (as a first approximation) steppe ancestry in Southeastern Europe from the Bronze Age onward was largely mediated by descendants of Yamnaya and local Balkan populations and not by earlier waves out of the steppe that affected the region sporadically. This admixture need not have taken place in one locality, as indicated by the presence of Yamnaya-like individuals in several regions of the Balkans, spatially beyond both the cultural transition
zone between steppe pastoralist and settled populations (32), and the geographical one from the Eastern European flatlands into mountainous areas.''
https://reich.hms.harvard.edu/sites/...ronzeAge_2.pdf
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