What are the Ainu like genetically?

[Sikeliot]

My guess is like Turkic people.

[Insuperable]

What Sardinians are to Europeans Ainu's are to East Asians. So, they are the most East Asian shifted people if I am not mistaken based on data I saw several years ago.

[Iloko]

Any opinions on this paper?: https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4701090/