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I checked Y-Full, the Illyrian samples from Croatia do not belong to the same subclade that predominates among contemporary Albanians:
https://www.yfull.com/tree/J-Y86930/
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K36 Similitude map for Iron Age Alderney (named Channel Islands in the paper, but all samples are from Alderney):
n=3 [samples: I16430, I26628, I16505]
Some top closest single distances (when using more of regional populations):
Spoiler!
And here are the K36 scores used (either there is a lot of noise, or this North African came with earliest Romans?):
Sample Alderney_IA
Amerindian 0
Arabian 0.20
Armenian 0
Basque 4.65
Central African 0.08
Central Euro 8.90
East African 0
East Asian 0
East Balkan 0.95
East Central Asian 0
East Central Euro 6.20
East Med 0
Eastern Euro 2.37
Fennoscandian 6.35
French 4.76
Iberian 21.69
Indo-Chinese 0
Italian 11.48
Malayan 0
Near Eastern 0
North African 1.91
North Atlantic 13.19
North Caucasian 0
North Sea 15.44
Northeast African 0
Oceanian 0.25
Omotic 0
Pygmy 0
Siberian 0
South Asian 0.24
South Central Asian 0
South Chinese 0
Volga-Ural 0.29
West African 0
West Caucasian 0.38
West Med 0.66
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If we assume that those IA Alderney people were representative of ancient Armoricans, then it seems that Normandy and East Brittany (Gallo-speaking zone, around Rennes) are more similar to ancient Armoricans, than people from West Brittany. Western Brittany (actual Breton-speaking areas) can probably be modeled as a mix of IA Alderney + IA Britain.
^^^
This G25 average for Channel Islands probably includes a low resolution sample I26629.
This sample should be removed and only the 3 decent quality samples should be used:
I16430, I26628, I16505
Also I think that you might get good fits if you model Bretons as a mix of Channel Islands IA + British IA.
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I think you underestimate the Anglo-Saxon/Germanic influence. In the Schiffels study they compared Anglo-Saxons to late Iron Age British (who look potentially Germanic mixed to me), where they came up with the figure of 38% 'Anglo-Saxon' for Eastern English, with a spread of 25-50%, and 30% even for Scotland and Wales. In the Viking study from last year the ancient Scandinavian-like input in England was estimated as 42%, with 37% being ancient-Danish like.
But who knows whether there was already Germanic admix in England in the Iron Age, I'm just speculating based on some things.
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The Gauls were descriibed by the Romans as northern-looking, so writing "Gallic/Iberian", as if they were equivalents, makes no sense. Check the K36 of Channel Islands Iron Age population (which is probably a good proxy for Armoricans). They were pretty much identical as modern North French, except for Breton-speaking part of Bretagne (remember that people in Eastern Bretagne speak Gallo, and they never spoke Breton in this part - around Rennes).
You were probably right that IA Britain is necessary to model Bretons, but it only applies to West Bretagne (Breton-speaking part).
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Well, what you called a "shift of England (between EIA and LIA) towards Germanics" also looks like a "shift of England (between EIA and LIA) towards Scotland_IA" to me. So this admixture could be from northern parts of Britain, rather than from areas across the North Sea. Another issue - you should check the regional distribution of England_EIA, England_MIA and England_LIA samples. Because for example if England_EIA samples are mainly from South England; England_MIA samples mainly from English Midlands; and England_LIA mainly from Northern England - then this observed "shift" is not a shift at all, but just a function of geography (because already in the Early Iron Age there was more of EEF admixture in the south of Britain than in the north).
The next thing:
What about uniparentals? If you are unsure about something based on auDNA, maybe a good idea is to check haplogroups?
Maciamo calculated haplogroup statistics and posted on his Eupedia Forum, I'm quoting him:
https://www.eupedia.com/forum/thread...l=1#post636856
Early Iron Age Britain (n=14)
I2a2-M223>Y3259>Y6098>PF692
R1b-P297
R1b-M269
R1b-L52
R1b-U106>S264
R1b-P312
R1b-P312
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13>Z251
R1b-P312>L21>DF13>Z251>A11676
R1b-P312>L21>DF13>DF21
R1b-P312>L21>DF13>DF49>DF23
R1b-P312>DF27>Z195>Z29704>S11475
Middle & Late Iron Age Britain (n=146)
F
F
F
F
G2a-L497>Z1815>Z1816>BY27899
G2a-L497>Z1815>Z1816>Z1823
G2a-L497>Z1815>Z1816>Z1823
G2a-L497>Z1815>Z1816>Z1823>Z726>S23438
G2a-L497>Z1815>Z1816>Z1823>Z726>Z16775
I1
I2a1a-M26>L160
I2a1a-M26>L160>Z105
I2a2a-M223>Y3259>Y6098
I2a2a-M223>Z284>L1195
I2a2a-M223>Z284>L1195
I2a2a-M223>Z284>L1195>Y3684
I2a2a-M223>Z284>L1195>Y3684
R1b-M269
R1b-M269
R1b-M269
R1b-M269
R1b-M269
R1b-M269
R1b-M269
R1b-M269
R1b-M269
R1b-M269>PF7562>Y83965
R1b-L52
R1b-L52
R1b-L52
R1b-L52>L151
R1b-L52>L151
R1b-L52>L151>S1200
R1b-L52>L151>P312
R1b-L52>L151>P312
R1b-L52>L151>P312
R1b-L52>L151>P312
R1b-L52>L151>P312
R1b-L52>L151>P312
R1b-L52>L151>P312
R1b-L52>L151>P312
R1b-L52>L151>P312
R1b-L52>L151>P312
R1b-L52>L151>P312>S461
R1b-L52>L151>P312>Z30597
R1b-L52>L151>P312>Z30597
R1b-P312>L21
R1b-P312>L21
R1b-P312>L21
R1b-P312>L21
R1b-P312>L21
R1b-P312>L21
R1b-P312>L21
R1b-P312>L21
R1b-P312>L21
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13
R1b-P312>L21>DF13>A1101
R1b-P312>L21>DF13>CTS1751>FGC49766
R1b-P312>L21>DF13>DF1
R1b-P312>L21>DF13>DF1>S5668
R1b-P312>L21>DF13>DF21
R1b-P312>L21>DF13>DF21
R1b-P312>L21>DF13>DF21>BY11117
R1b-P312>L21>DF13>DF21>DF25>DF5
R1b-P312>L21>DF13>DF21>DF25>Y12651
R1b-P312>L21>DF13>DF21>S5488
R1b-P312>L21>DF13>DF21>S5488
R1b-P312>L21>DF13>DF21>S5488
R1b-P312>L21>DF13>DF21>Y2890
R1b-P312>L21>DF13>DF49
R1b-P312>L21>DF13>DF49
R1b-P312>L21>DF13>DF49
R1b-P312>L21>DF13>DF49
R1b-P312>L21>DF13>DF49>DF23
R1b-P312>L21>DF13>DF49>DF23>Z2961
R1b-P312>L21>DF13>DF49>BY23924
R1b-P312>L21>DF13>FGC5494
R1b-P312>L21>DF13>FGC5494>S1088
R1b-P312>L21>DF13>FGC5494>S1088
R1b-P312>L21>DF13>FGC5494>S7958>BY33481
R1b-P312>L21>DF13>FGC5494>Y31855
R1b-P312>L21>DF13>S1026>BY173541
R1b-P312>L21>DF13>S1026>Z16887
R1b-P312>L21>DF13>S1051>FGC19428
R1b-P312>L21>DF13>Y14049>BY9003
R1b-P312>L21>DF13>Z251
R1b-P312>L21>DF13>Z251
R1b-P312>L21>DF13>Z251
R1b-P312>L21>DF13>Z251
R1b-P312>L21>DF13>Z251
R1b-P312>L21>DF13>Z251
R1b-P312>L21>DF13>Z251>Y11273
R1b-P312>L21>DF13>Z253
R1b-P312>L21>DF13>Z253
R1b-P312>L21>DF13>Z253
R1b-P312>L21>DF13>Z253>Z2185
R1b-P312>L21>DF13>Z253>Z2185>Z2186
R1b-P312>L21>DF13>Z253>Z2534
R1b-P312>L21>DF13>Z253>Z2534>A14
R1b-P312>L21>DF13>Z253>Z2534>BY13089
R1b-P312>L21>DF13>Z255>Z16436
R1b-P312>L21>DF63
R1b-P312>L21>DF63
R1b-P312>L21>DF63
R1b-P312>L21>DF63
R1b-P312>L21>DF63
R1b-P312>L21>DF63
R1b-P312>L21>DF63>CTS6919
R1b-P312>L21>DF63>FGC36421
R1b-P312>L238>Z2245
R1b-P312>DF27
R1b-P312>DF27>FT318890>Y3267
R1b-P312>DF27>Y14529
R1b-P312>DF27>Y30754
R1b-P312>DF27>Y30754>Y85515
R1b-P312>DF27>Z195>Z262
R1b-P312>DF27>Z225>Y89302
R1b-P312>DF27>Z2572
R1b-P312>U152>L2
R1b-P312>U152>L2
R1b-P312>U152>L2>Y3961>Y17997
R1b-P312>U152>L2>Z49>Z142>S7402
R1b-P312>U152>L2>Z49>Z142>S7402>FGC12384
R1b-P312>U152>L2>Z49>Z142>Z51>Z57>Z53
Haplogroup frequency:
R1b = 88.3% (including 58.2% of L21, 6.8% of M269, 5.5% of DF27, 4.1% of L2 and 0% of U106)
G2a-L497 = 3.4%
I2a2a-M223 = 3.4%
F = 2.7%
I2a1a-M26 = 1.4%
I1 = 0.7%
=====
In other words, ABSOLUTE LACK of Germanic Y-DNA haplogroups in Iron Age Britain, except for maybe one singleton I1.
What percentage of English males today carry Germanic Y-DNA ??? I don't remember exactly, but surely double-digits.
And in IA England it was no more than 1% (if that singleton I1 is real - and not misdated or a wrongly assigned haplo).
To sum up, there was no male-mediated Germanic admixture in Iron Age England. Maybe female-mediated.
Maybe Britons were buying blond-haired Germanic slave girls and got Germanic admixture - if you want to believe this.
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